The topography and geology of the Fayum province of Egypt

by H. J. L. Beadnell

Plate XII.

RAISED BEACH UNCONFORMABLY OVERLYING MIDDLE EOCENE LIMESTONES (BIRKET EL QURUN SERIES) IN THE DESERT EAST OF SIRSENA.

In this connection it is interesting to notice the observation of so eminent a palæontologist as Prof. H. F. Osborn. In two recent addresses[83] to the New York Academy of Sciences he pertinently points out his belief that the African continent has been a great centre of radiation of certain groups of the mammalia, and especially mentions the Proboscidea as likely to have been evolved in the Ethiopian region. Our discoveries in the Fayûm and Andrews’s determinations, made subsequently to these addresses, so completely confirm this view, at any rate with regard to the elephants, that it may not be out of place to give here a somewhat lengthy extract of his “Theory of Successive Invasions of an African Fauna into Europe” (op. cit. pp. 56-58). “In Europe there are in the Upper Eocene two classes of animals, first those which have their ancestors in the older rocks; second, the class including certain highly specialized animals which have no ancestors in the older rocks, among these, perhaps, are the peculiar flying rodents or Anomaluridæ, now confined to Africa, and secondly the highly specialized even-toed ruminant types the anoplotheres, xiphodonts and others, the discovery of which in the gypse near Paris Cuvier has made famous. It is tempting to imagine that these animals did not evolve in Europe but that they represent what may be called the first invasion of Europe by African types from the Ethiopian region.

“It is a curious fact that the African continent as a great theater of adaptive radiation of Mammalia has not been sufficiently considered. It is true that it is the dark continent of palæontology for it has no fossil mammal history; but it by no means follows that the Mammalia did not enjoy there an extensive evolution.[84]

“Although it is quite probable that this idea has been advanced before, most writers speak mainly or exclusively of the invasion of Africa by European types. Blanford and Allen, it is true, have especially dwelt upon the likeness of the Oriental and Ethiopian fauna, but not in connection with its antecedent cause. This cause I believe to have been mainly an invasion from south to north, correlated with the northern extension of Ethiopian climate and flora during the Middle Tertiary. It is in a less measure due to a migration from north to south. Let us therefore clearly set forth the hypothesis of the Ethiopian region or South Africa as a great center of independent evolution and as the source of successive northward migrations of animals, some of which ultimately reached even the extremity of South America, I refer to the Mastodons. This hypothesis is clearly implied if not stated by Blanford in 1876 in his paper upon the African element in the fauna of India.

“The first of these migrations we may suppose brought in certain highly specialized ruminants of the Upper Eocene, the anomalures or peculiar flying rodents of Africa; with this invasion may have come the pangolins and ard varks, and possibly certain armadillos, Dasypodidæ, if M. Filhol’s identification of Necrodasypus is correct. A second invasion of great distinctness may be that which marks the beginning of the Miocene when the mastodons and dinotheres first appear in Europe, also the earliest of the antelopes. A third invasion may be represented in the base of the Pliocene by the increasing number of antelopes, the great giraffes of the Ægean plateau and in the upper Pliocene by the hippopotami. With these forms came the rhinoceroses with no incisor or cutting teeth, similar to the smaller African rhinoceros, R. bicornis. Another recently discovered African immigrant upon the Island of Samos in the Ægean plateau is Pliohyrax or Leptodon, a very large member of the Hyracoidea, probably aquatic in its habits, indicating that this order enjoyed an extensive adaptive radiation in Tertiary times.

“It thus appears that the Proboscidea, Hyracoidea, certain edentata, the antelopes, the giraffes, the hippopotami, the most specialized ruminants, and among the rodents, the anomalures, dormice and jerboas, among monkeys the baboons, may all have enjoyed their original adaptative radiation in Africa; that they survived after the glacial period, only in the Oriental or Indo-Malayan region, and that this accounts for the marked community of fauna between this region and the Ethiopian as observed by Blanford and Allen.

“Against the prevalent theory of Oriental origin of these animals are: first, the fact observed by Blanford and Lydekker in the Bugti Beds (Sind) that the Oligocene or lower Miocene fauna of the Orient is markedly European in type; second, that if these animals had originated in Asia some of them would have found their way to North America; third, the fact that all these animals appear suddenly and without any known ancestors in older geological formations. These are the main facts in favor of the Ethiopian migration hypothesis.”

That Professor Osborn’s main contention has already been partly proved by the Fayûm mammal discoveries is apparent, and how far his detailed remarks are confirmed will be seen when the new fauna has been more completely explored and examined.