This pretty creature inhabits most of the warm portions of the continental Indian region, from India and Ceylon to South China and Malacca. The back is a rich dark brown, divided from the yellow of the head by a narrow black line which extends from eye to eye and forwards to each nostril. A conspicuous yellow band runs from the eyes to the hind-limbs. The sides of the body and the limbs are mottled yellow and brown. The under parts are dirty buff; the throat of the male is black. The intensity of colouring varies individually and from time to time, the contrast between the brown and yellow being occasionally very brilliant. Total length up to 3 inches, the male being the smaller sex.
"I have been told by both English and natives that this frog was unknown in Singapore until some nine or ten years ago, when it was introduced by a half-caste (why, it is not known), and that it rapidly spread about the island. It is now well known as the 'Bullfrog' by the English in Singapore, and detested for the noise it makes at night. The voice of these rotund animals can be heard every night after heavy rain; it is a deep guttural croak, 'wau-auhhhh,' very strident and prolonged. The males croak while floating on the surface of the water, the single vocal sac under the mouth inflated like a globe, and the arms and legs extended. They can hop well on land and are good swimmers. The skin is excessively slimy; the secretion comes off profusely, and dries on the hand into a sort of white gum, with a faint aromatic smell. This gum dissolves in hot water and coagulates in cold. The general appearance of these frogs is very stout, their girth being about twice the length from snout to vent. The tongue, which is oblong in spirit specimens, in life is very elastic, assuming, when extended, a vermiform shape and reaching about 4 cm. in length. They appear after sunset, crawling on old wood and feeding on white ants."
Sub-Fam. 2. Dyscophinae.–With teeth in the upper jaw.
This small group of nine genera, with scarcely more than one dozen species, all with one exception living in Madagascar, has been separated by Boulenger from the Engystomatinae merely on account of the presence of teeth on the upper jaw and on the vomerine margin of the palatine bones. He himself remarks that Calluella may be considered a toothed Hypopachus, and Plethodontohyla a toothed Callula. These are obvious cases of convergent analogy. Except for the teeth, the Indian Calluella would be merged into the American Hypopachus, and this would present an instance of the most puzzling geographical distribution. In the case of the other two genera, one Indian and Malayan, the other Malagasy, no such suspicion would arise, since there are many other instances of such a coincidence of distribution. There is the same divergence or unsettled condition in the modification of various parts in the Dyscophinae as in the Engystomatinae. The precoracoid bars are weak and curved backwards, and closely pressed against the strong coracoids, in Dyscophus, Calluella and Platypelis, while these elements are reduced to unossified bars, and the clavicular portions completely lost, in Plethodontohyla and in Phrynocara. The omosternum is absent and the metasternum is small in all except Dyscophus, in which both these parts are exceptionally well developed and large, although remaining unossified. The palate of Dyscophus and Calluella is provided with curious, serrated dermal folds like those which are so common in the Engystomatinae; and well-developed discs on the fingers and toes, supported by T-shaped phalanges, are possessed by Platypelis, Cophyla and others. The sacral diapophyses are dilated. The pupil is either horizontal or vertical. Those which are provided with discs to the fingers and toes are climbers, and mostly slender and long-legged, sometimes of very small size, for instance Cophyla, the body of which is scarcely one inch in length.
The genera can be determined by means of the following key:–[[103]]
A. Pupil vertical. Palatine teeth in long transverse series.
a. Precoracoids ossified. Tips of fingers and toes not dilated.
Sternum very large. Madagascar .......... Dyscophus.
Sternum small. Burmah .......... Calluella.
b. Precoracoids not ossified. Tips dilated .......... Plethodontohyla.