Metagonimus, Katsurada, 1913; Yokogawa, Leiper, 1913.

Resembles in general structure Heterophyes. In the arrangement of its ventral genital suckers resembles but differs from that of Tocotrema,[273] Looss. The ventral and genital suckers lie laterally and on the right.

Metagonimus yokogawai. Katsurada, 1913.

Syn.: Yokogawa yokogawai, Leiper, 1913.

Fig. 164.—Metagonimus yokogawai, Katsurada, 1913: the spines are only shown over a small part of the skin. (After Leiper.)

One to 1·5 mm. long, seldom 2·5 mm., and 0·4 to 0·7 mm. broad; elliptical in shape. The body is thickly covered with nail-shaped spines about 10 µ long. Oral sucker 77 µ, to 85 µ in diameter. Ventral sucker characteristic and peculiar 0·12 to 0·14 mm. by 0·08 to 1 mm. It is a sac-like organ placed deeply in the body, but does not open as in other flukes on the ventral surface. Testes elliptical, not quite symmetrically placed at the hind end of the body. Vesicula seminalis retort-shaped, situated transversely, internal to the ventral sucker. Pars prostatica present. Ejaculatory duct opens with the uterus into a genital sinus, which, together with the internal opening of the ventral sucker, opens into a pit at the front of the ventral sucker. The opening of the genital sinus and that of the ventral sucker are furnished with a complex muscular apparatus. Ovary spherical, 0·12 to 0·13 mm. in diameter, lies in the middle of the hind body. Receptaculum seminis and Laurer’s canal present. Vitellaria in the hind half of the body, consisting of about ten acini on each side. Shell gland to the left of the ovary. Uterus forms three to four transverse coils. Eggs elliptical, double contoured, yellowish-brown in colour. There is no shoulder below the operculum as in the eggs of Cl. sinensis. At the rounder end there is a thickening or knob different from the spine-like or hook-like process seen in Cl. sinensis. Dimensions 28 µ by 16 µ.

Habitat.—Mainly in upper or middle portion of jejunum, rarely in cæcum. They penetrate deep into the mucosa, but not into the submucosa, and post mortem appear as a number of small brown points. They frequently occur in the solitary glands, which they destroy. They cause chronic catarrh of the gut. Parasitic in man and mammals.

Geographical Distribution.—Japan.