That adaptation to environment is the necessary condition of evolution we do not question for a moment. It is quite evident that a species would disappear, should it fail to bend to the conditions of existence which are imposed on it. But it is one thing to recognize that outer circumstances are forces evolution must reckon with, another to claim that they are the directing causes of evolution. This latter theory is that of mechanism. It excludes absolutely the hypothesis of an original impetus, I mean an internal push that has carried life, by more and more complex forms, to higher and higher destinies. Yet this impetus is evident, and a mere glance at fossil species shows us that life need not have evolved at all, or might have evolved only in very restricted limits, if it had chosen the alternative, much more convenient to itself, of becoming anchylosed in its primitive forms. Certain Foraminifera have not varied since the Silurian epoch. Unmoved witnesses of the innumerable revolutions that have upheaved our planet, the Lingulae are to-day what they were at the remotest times of the paleozoic era.
The truth is that adaptation explains the sinuosities of the movement of evolution, but not its general directions, still less the movement itself.[51] The road that leads to the town is obliged to follow the ups and downs of the hills; it adapts itself to the accidents of the ground; but the accidents of the ground are not the cause of the road, nor have they given it its direction. At every moment they furnish it with what is indispensable, namely, the soil on which it lies; but if we consider the whole of the road, instead of each of its parts, the accidents of the ground appear only as impediments or causes of delay, for the road aims simply at the town and would fain be a straight line. Just so as regards the evolution of life and the circumstances through which it passes—with this difference, that evolution does not mark out a solitary route, that it takes directions without aiming at ends, and that it remains inventive even in its adaptations.
But, if the evolution of life is something other than a series of adaptations to accidental circumstances, so also it is not the realization of a plan. A plan is given in advance. It is represented, or at least representable, before its realization. The complete execution of it may be put off to a distant future, or even indefinitely; but the idea is none the less formulable at the present time, in terms actually given. If, on the contrary, evolution is a creation unceasingly renewed, it creates, as it goes on, not only the forms of life, but the ideas that will enable the intellect to understand it, the terms which will serve to express it. That is to say that its future overflows its present, and can not be sketched out therein in an idea.
There is the first error of finalism. It involves another, yet more serious.
If life realizes a plan, it ought to manifest a greater harmony the further it advances, just as the house shows better and better the idea of the architect as stone is set upon stone. If, on the contrary, the unity of life is to be found solely in the impetus that pushes it along the road of time, the harmony is not in front, but behind. The unity is derived from a vis a tergo: it is given at the start as an impulsion, not placed at the end as an attraction. In communicating itself, the impetus splits up more and more. Life, in proportion to its progress, is scattered in manifestations which undoubtedly owe to their common origin the fact that they are complementary to each other in certain aspects, but which are none the less mutually incompatible and antagonistic. So the discord between species will go on increasing. Indeed, we have as yet only indicated the essential cause of it. We have supposed, for the sake of simplicity, that each species received the impulsion in order to pass it on to others, and that, in every direction in which life evolves, the propagation is in a straight line. But, as a matter of fact, there are species which are arrested; there are some that retrogress. Evolution is not only a movement forward; in many cases we observe a marking-time, and still more often a deviation or turning back. It must be so, as we shall show further on, and the same causes that divide the evolution movement often cause life to be diverted from itself, hypnotized by the form it has just brought forth. Thence results an increasing disorder. No doubt there is progress, if progress mean a continual advance in the general direction determined by a first impulsion; but this progress is accomplished only on the two or three great lines of evolution on which forms ever more and more complex, ever more and more high, appear; between these lines run a crowd of minor paths in which, on the contrary, deviations, arrests, and set-backs, are multiplied. The philosopher, who begins by laying down as a principle that each detail is connected with some general plan of the whole, goes from one disappointment to another as soon as he comes to examine the facts; and, as he had put everything in the same rank, he finds that, as the result of not allowing for accident, he must regard everything as accidental. For accident, then, an allowance must first be made, and a very liberal allowance. We must recognize that all is not coherent in nature. By so doing, we shall be led to ascertain the centres around which the incoherence crystallizes. This crystallization itself will clarify the rest; the main directions will appear, in which life is moving whilst developing the original impulse. True, we shall not witness the detailed accomplishment of a plan. Nature is more and better than a plan in course of realization. A plan is a term assigned to a labor: it closes the future whose form it indicates. Before the evolution of life, on the contrary, the portals of the future remain wide open. It is a creation that goes on for ever in virtue of an initial movement. This movement constitutes the unity of the organized world—a prolific unity, of an infinite richness, superior to any that the intellect could dream of, for the intellect is only one of its aspects or products.
But it is easier to define the method than to apply it. The complete interpretation of the evolution movement in the past, as we conceive it, would be possible only if the history of the development of the organized world were entirely known. Such is far from being the case. The genealogies proposed for the different species are generally questionable. They vary with their authors, with the theoretic views inspiring them, and raise discussions to which the present state of science does not admit of a final settlement. But a comparison of the different solutions shows that the controversy bears less on the main lines of the movement than on matters of detail; and so, by following the main lines as closely as possible, we shall be sure of not going astray. Moreover, they alone are important to us; for we do not aim, like the naturalist, at finding the order of succession of different species, but only at defining the principal directions of their evolution. And not all of these directions have the same interest for us: what concerns us particularly is the path that leads to man. We shall therefore not lose sight of the fact, in following one direction and another, that our main business is to determine the relation of man to the animal kingdom, and the place of the animal kingdom itself in the organized world as a whole.
To begin with the second point, let us say that no definite characteristic distinguishes the plant from the animal. Attempts to define the two kingdoms strictly have always come to naught. There is not a single property of vegetable life that is not found, in some degree, in certain animals; not a single characteristic feature of the animal that has not been seen in certain species or at certain moments in the vegetable world. Naturally, therefore, biologists enamored of clean-cut concepts have regarded the distinction between the two kingdoms as artificial. They would be right, if definition in this case must be made, as in the mathematical and physical sciences, according to certain statical attributes which belong to the object defined and are not found in any other. Very different, in our opinion, is the kind of definition which befits the sciences of life. There is no manifestation of life which does not contain, in a rudimentary state—either latent or potential,—the essential characters of most other manifestations. The difference is in the proportions. But this very difference of proportion will suffice to define the group, if we can establish that it is not accidental, and that the group as it evolves, tends more and more to emphasize these particular characters. In a word, the group must not be defined by the possession of certain characters, but by its tendency to emphasize them. From this point of view, taking tendencies rather than states into account, we find that vegetables and animals may be precisely defined and distinguished, and that they correspond to two divergent developments of life.
This divergence is shown, first, in the method of alimentation. We know that the vegetable derives directly from the air and water and soil the elements necessary to maintain life, especially carbon and nitrogen, which it takes in mineral form. The animal, on the contrary, cannot assimilate these elements unless they have already been fixed for it in organic substances by plants, or by animals which directly or indirectly owe them to plants; so that ultimately the vegetable nourishes the animal. True, this law allows of many exceptions among vegetables. We do not hesitate to class amongst vegetables the Drosera, the Dionaea, the Pinguicula, which are insectivorous plants. On the other hand, the fungi, which occupy so considerable a place in the vegetable world, feed like animals: whether they are ferments, saprophytes or parasites, it is to already formed organic substances that they owe their nourishment. It is therefore impossible to draw from this difference any static definition such as would automatically settle in any particular case the question whether we are dealing with a plant or an animal. But the difference may provide the beginning of a dynamic definition of the two kingdoms, in that it marks the two divergent directions in which vegetables and animals have taken their course. It is a remarkable fact that the fungi, which nature has spread all over the earth in such extraordinary profusion, have not been able to evolve. Organically they do not rise above tissues which, in the higher vegetables, are formed in the embryonic sac of the ovary, and precede the germinative development of the new individual.[52] They might be called the abortive children of the vegetable world. Their different species are like so many blind alleys, as if, by renouncing the mode of alimentation customary amongst vegetables, they had been brought to a standstill on the highway of vegetable evolution. As to the Drosera, the Dionaea, and insectivorous plants in general, they are fed by their roots, like other plants; they too fix, by their green parts, the carbon of the carbonic acid in the atmosphere. Their faculty of capturing, absorbing and digesting insects must have arisen late, in quite exceptional cases where the soil was too poor to furnish sufficient nourishment. In a general way, then, if we attach less importance to the presence of special characters than to their tendency to develop, and if we regard as essential that tendency along which evolution has been able to continue indefinitely, we may say that vegetables are distinguished from animals by their power of creating organic matter out of mineral elements which they draw directly from the air and earth and water. But now we come to another difference, deeper than this, though not unconnected with it.