We will not dwell here on a point we have treated at length in a former work. Let us merely recall that the progress of the nervous system has been effected both in the direction of a more precise adaptation of movements and in that of a greater latitude left to the living being to choose between them. These two tendencies may appear antagonistic, and indeed they are so; but a nervous chain, even in its most rudimentary form, successfully reconciles them. On the one hand, it marks a well-defined track between one point of the periphery and another, the one sensory, the other motor. It has therefore canalized an activity which was originally diffused in the protoplasmic mass. But, on the other hand, the elements that compose it are probably discontinuous; at any rate, even supposing they anastomose, they exhibit a functional discontinuity, for each of them ends in a kind of cross-road where probably the nervous current may choose its course. From the humblest Monera to the best endowed insects, and up to the most intelligent vertebrates, the progress realized has been above all a progress of the nervous system, coupled at every stage with all the new constructions and complications of mechanism that this progress required. As we foreshadowed in the beginning of this work, the rôle of life is to insert some indetermination into matter. Indeterminate, i.e. unforeseeable, are the forms it creates in the course of its evolution. More and more indeterminate also, more and more free, is the activity to which these forms serve as the vehicle. A nervous system, with neurones placed end to end in such wise that, at the extremity of each, manifold ways open in which manifold questions present themselves, is a veritable reservoir of indetermination. That the main energy of the vital impulse has been spent in creating apparatus of this kind is, we believe, what a glance over the organized world as a whole easily shows. But concerning the vital impulse itself a few explanations are necessary.
It must not be forgotten that the force which is evolving throughout the organized world is a limited force, which is always seeking to transcend itself and always remains inadequate to the work it would fain produce. The errors and puerilities of radical finalism are due to the misapprehension of this point. It has represented the whole of the living world as a construction, and a construction analogous to a human work. All the pieces have been arranged with a view to the best possible functioning of the machine. Each species has its reason for existence, its part to play, its allotted place; and all join together, as it were, in a musical concert, wherein the seeming discords are really meant to bring out a fundamental harmony. In short, all goes on in nature as in the works of human genius, where, though the result may be trifling, there is at least perfect adequacy between the object made and the work of making it.
Nothing of the kind in the evolution of life. There, the disproportion is striking between the work and the result. From the bottom to the top of the organized world we do indeed find one great effort; but most often this effort turns short, sometimes paralyzed by contrary forces, sometimes diverted from what it should do by what it does, absorbed by the form it is engaged in taking, hypnotized by it as by a mirror. Even in its most perfect works, though it seems to have triumphed over external resistances and also over its own, it is at the mercy of the materiality which it has had to assume. It is what each of us may experience in himself. Our freedom, in the very movements by which it is affirmed, creates the growing habits that will stifle it if it fails to renew itself by a constant effort: it is dogged by automatism. The most living thought becomes frigid in the formula that expresses it. The word turns against the idea.
The letter kills the spirit. And our most ardent enthusiasm, as soon as it is externalized into action, is so naturally congealed into the cold calculation of interest or vanity, the one takes so easily the shape of the other, that we might confuse them together, doubt our own sincerity, deny goodness and love, if we did not know that the dead retain for a time the features of the living.
The profound cause of this discordance lies in an irremediable difference of rhythm. Life in general is mobility itself; particular manifestations of life accept this mobility reluctantly, and constantly lag behind. It is always going ahead; they want to mark time. Evolution in general would fain go on in a straight line; each special evolution is a kind of circle. Like eddies of dust raised by the wind as it passes, the living turn upon themselves, borne up by the great blast of life. They are therefore relatively stable, and counterfeit immobility so well that we treat each of them as a thing rather than as a progress, forgetting that the very permanence of their form is only the outline of a movement. At times, however, in a fleeting vision, the invisible breath that bears them is materialized before our eyes. We have this sudden illumination before certain forms of maternal love, so striking, and in most animals so touching, observable even in the solicitude of the plant for its seed. This love, in which some have seen the great mystery of life, may possibly deliver us life's secret. It shows us each generation leaning over the generation that shall follow. It allows us a glimpse of the fact that the living being is above all a thoroughfare, and that the essence of life is in the movement by which life is transmitted.
This contrast between life in general, and the forms in which it is manifested, has everywhere the same character. It might be said that life tends toward the utmost possible action, but that each species prefers to contribute the slightest possible effort. Regarded in what constitutes its true essence, namely, as a transition from species to species, life is a continually growing action. But each of the species, through which life passes, aims only at its own convenience. It goes for that which demands the least labor. Absorbed in the form it is about to take, it falls into a partial sleep, in which it ignores almost all the rest of life; it fashions itself so as to take the greatest possible advantage of its immediate environment with the least possible trouble. Accordingly, the act by which life goes forward to the creation of a new form, and the act by which this form is shaped, are two different and often antagonistic movements. The first is continuous with the second, but cannot continue in it without being drawn aside from its direction, as would happen to a man leaping, if, in order to clear the obstacle, he had to turn his eyes from it and look at himself all the while.
Living forms are, by their very definition, forms that are able to live. In whatever way the adaptation of the organism to its circumstances is explained, it has necessarily been sufficient, since the species has subsisted. In this sense, each of the successive species that paleontology and zoology describes was a success carried off by life. But we get a very different impression when we refer each species to the movement that has left it behind on its way, instead of to the conditions into which it has been set. Often this movement has turned aside; very often, too, it has stopped short; what was to have been a thoroughfare has become a terminus. From this new point of view, failure seems the rule, success exceptional and always imperfect. We shall see that, of the four main directions along which animal life bent its course, two have led to blind alleys, and, in the other two, the effort has generally been out of proportion to the result.
Documents are lacking to reconstruct this history in detail, but we can make out its main lines. We have already said that animals and vegetables must have separated soon from their common stock, the vegetable falling asleep in immobility, the animal, on the contrary, becoming more and more awake and marching on to the conquest of a nervous system. Probably the effort of the animal kingdom resulted in creating organisms still very simple, but endowed with a certain freedom of action, and, above all, with a shape so undecided that it could lend itself to any future determination. These animals may have resembled some of our worms, but with this difference, however, that the worms living to-day, to which they could be compared, are but the empty and fixed examples of infinitely plastic forms, pregnant with an unlimited future, the common stock of the echinoderms, molluscs, arthropods, and vertebrates.
One danger lay in wait for them, one obstacle which might have stopped the soaring course of animal life. There is one peculiarity with which we cannot help being struck when glancing over the fauna of primitive times, namely, the imprisonment of the animal in a more or less solid sheath, which must have obstructed and often even paralyzed its movements. The molluscs of that time had a shell more universally than those of to-day. The arthropods in general were provided with a carapace; most of them were crustaceans. The more ancient fishes had a bony sheath of extreme hardness.[59] The explanation of this general fact should be sought, we believe, in a tendency of soft organisms to defend themselves against one another by making themselves, as far as possible, undevourable. Each species, in the act by which it comes into being, trends towards that which is most expedient. Just as among primitive organisms there were some that turned towards animal life by refusing to manufacture organic out of inorganic material and taking organic substances ready made from organisms that had turned toward the vegetative life, so, among the animal species themselves, many contrived to live at the expense of other animals. For an organism that is animal, that is to say mobile, can avail itself of its mobility to go in search of defenseless animals, and feed on them quite as well as on vegetables. So, the more species became mobile, the more they became voracious and dangerous to one another. Hence a sudden arrest of the entire animal world in its progress towards higher and higher mobility; for the hard and calcareous skin of the echinoderm, the shell of the mollusc, the carapace of the crustacean and the ganoid breast-plate of the ancient fishes probably all originated in a common effort of the animal species to protect themselves against hostile species. But this breast-plate, behind which the animal took shelter, constrained it in its movements and sometimes fixed it in one place. If the vegetable renounced consciousness in wrapping itself in a cellulose membrane, the animal that shut itself up in a citadel or in armor condemned itself to a partial slumber. In this torpor the echinoderms and even the molluscs live to-day. Probably arthropods and vertebrates were threatened with it too. They escaped, however, and to this fortunate circumstance is due the expansion of the highest forms of life.