It follows from the above, that the Lower and Upper Cretaceous rocks are, from a botanical point of view, sharply separated from one another. The Palæozoic period, as we have seen, is characterised by the prevalance of "Flowerless" plants (Cryptogams), its higher vegetation consisting almost exclusively of Conifers. The Mesozoic period, as a whole, is characterised by the prevalence of the Cryptogamic group of the Ferns, and the Gymnospermic groups of the Conifers and the Cycads. Up to the close of the Lower Cretaceous, no Angiospermous Exogens are certainly known to have existed, and Monocotyledonous plants or Endogens are very poorly represented. With the Upper Cretaceous, however, a new era of plant-life, of which our present is but the culmination, commenced, with a great and apparently sudden development of new forms. In place of the Ferns, Cycads, and Conifers of the earlier Mesozoic deposits, we have now an astonishingly large number of true Angiospermous Exogens, many of them belonging to existing types; and along with these are various Monocotyledonous plants, including the first examples of the great and important group of the Palms. It is thus a matter of interest to reflect that plants closely related to those now
Fig. 186.—Cretaceous Angiosperms. a. Sassafras Cretaceum; b, Liriodendron Meekii; c, Leguminosites Marcouanus; d, Salix Meekii. (After Dana.) inhabiting the earth, were in existence at a time when the ocean was tenanted by Ammonites and Belemnites, and when land and sea and air were peopled by the extraordinary extinct Reptiles of the Mesozoic period.
As regards animal life, the Protozoans of the Cretaceous period are exceedingly numerous, and are represented by Foraminifera and Sponges. As we have already seen, the White Chalk itself is a deep-sea deposit, almost entirely composed of the microscopic shells of Foraminifers, along with Sponge-spicules, and organic débris of different kinds (see fig. 7). The green grains which are so abundant in several minor subdivisions of the Cretaceous, are also in many instances really casts in glauconite of the chambered shells of these minute organisms. A great many species of Foraminifera have been recognised in the Chalk; but the three principal genera are Globigerina, Rotalia (fig. 187), and Textularia—groups which are likewise characteristic of the "ooze" of the Atlantic and
Fig. 187—Kotalia Boueana. Pacific Oceans at great depths. The flints of the Chalk also commonly contain the shells of Foraminifera. The Upper Greensand has yielded in considerable numbers the huge Foraminifera described by Dr Carpenter under the name of Parkeria, the spherical shells of which are composed of sand-grains agglutinated together, and sometimes attain a diameter of two and a quarter inches. The Cretaceous Sponges are extremely numerous, and occur under a great number of varieties of shape and structure; but the two most characteristic genera are Siphonia and Ventriculites, both of which are exclusively confined to strata of this age. The Siphoniœ (fig. 188) consist of a pear-shaped, sometimes lobed head, supported by a longer or shorter stern, which breaks up at its base into a number of root-like processes of attachment. The water gained access to the interior of the Sponge by a number of minute openings covering the surface, and ultimately escaped by a single, large, chimney-shaped aperture at the summit. In some respects these sponges present a singular resemblance to the beautiful "Vitreous Sponges" (Holtenia or Pheronema) of the deep Atlantic; and, like these, they were probably denizens of a deep sea, The Ventriculites of the Chalk (fig. 189) is, however, a genus still more closely allied to the wonderful flinty Sponges, which have been shown, by the researches of the Porcupine, Lightning, and Challenger expeditions, to live half buried in the Calcareous ooze of the abysses of our great oceans. Many forms of this genus are known, having "usually the form of graceful vases, tubes, or funnels, variously ridged or grooved, or otherwise ornamented on the surface, frequently expanded above into a cup-like lip, and continued below into a bundle of fibrous roots. The minute structure of these bodies shows an extremely delicate tracery of fine tubes, sometimes empty, sometimes filled with loose calcareous matter dyed with peroxide of iron."—(Sir Wyville Thomson.) Many of the Chalk sponges, originally calcareous, have been converted into flint subsequently; but the Ventriculites are really composed
Fig. 188.—Siphonia ficus. Upper Greensand. Europe.
Fig. 189.—Ventriculites simplex. White Chalk. Britain. of this substance, and are therefore genuine "Siliceous Sponges," like the existing Venus's Flower-Basket (Euplectella). Like the latter, the skeleton was doubtless originally composed, in the young state, of disconnected six-rayed spicules, which ultimately become fixed together to constitute a continuous frame-work. The sea-water, as in the recent forms, must have been admitted to the interior of the Sponge by numerous apertures on its exterior, subsequently escaping by a single large opening at its summit.