In the complete scheme adopted by most naturalists, five categories include the evidences bearing upon the fact of evolution. These are Classification; Comparative Anatomy, or Morphology; Comparative Development, or _Embryology; Palæontology, which comprises the facts provided by fossil relics of animals and plants of earlier geological ages; and Geographical Distribution. Each of these divisions includes a descriptive and analytical series of facts, whose characteristics are "explained" or summarized in the form of the general principles of the respective divisions. Such principles, taken singly and collectively, constitute the evidences of evolution.
The particular nature of any one of these categories, evolved in the development of science practically in the order stated, depends upon the special quality of an animal which it selects for comparison and organization in connection with other similar facts, and also in its own mode of viewing its facts. One and the same organism may present materials for two, three, or even all five of these divisions, for they are by no means mutually exclusive. For example, a common cat possesses certain definite characteristics which give it a particular place when animals more or less like it are grouped or classified according to their degrees of resemblance and difference, in small genera of very similar forms, in larger tribes or orders of similar genera, and in more and more inclusive groups of these lesser divisions, such as the classes and phyla, or main branches of the animal tree. The common cat and its relatives are even earlier to be regarded as anatomical subjects, and their thorough analysis belongs to comparative anatomy,—a name which explains itself. The purpose of this department of natural history is to explore the entire range of animal forms and animal structures, and to determine the degree of resemblance and difference exhibited by the general characters of entire organisms and by the special qualities of their several systems of organs. It provides the data from which classification selects those which indicate mutual affinities with greatest precision and surety. But its materials are all the facts of animal structure, and because each and every known organism can be and must be studied, the investigator engaged in formulating the evidence of evolution has at his disposal all the data referring to the entire realm of animals. The data of embryology are likewise coextensive with the territory of the animal world, for we do not know of any form which does not change in the course of its life history. An adult cat is the product of a kitten which is itself the result of a long series of changes from earlier and simpler conditions. In so far as it deals with structures in the making, embryology is a study of anatomy, but as it is concerned primarily with all of the plastic remodeling which animals undergo during the production of their final forms, it is an independent study. Nevertheless we shall learn how intimate are the relations of these two divisions of zoölogy and how the evolutionary teachings of each body of fact support and supplement those of the other.
Palæontology searches everywhere among the deposits of earlier ages for links to be fitted into their proper sequence of time, from which it constructs the chain of diverse types leading down to the species of the present. A cat of to-day is therefore viewed in an entirely different connection, as the last term in a consecutive series of species. Forming alliances with geology, and even with physics and chemistry, this department of zoölogy endeavors to reconstruct the past from what it learns to-day about organisms and the conditions under which they live. Finally the observations that cats of various kinds do not occur everywhere in the world, but only in certain more or less restricted localities, belong to the subject of geographical distribution, and illustrate its nature.
Our task is to learn the teachings of these several divisions by recalling and putting together what we know already about the commonest animals, or noting what can be observed in a visit to a zoölogical garden and aquarium. On account of the present limitations of time, the subject of classification will be combined with comparative anatomy; embryology will be taken up together with these subjects; palæontology will be the main subject of the next discussion, which will include also a brief statement of the meaning of distribution. Then we will be prepared to study nature to see how evolution works.
II
THE STRUCTURE AND DEVELOPMENT OF ANIMALS AS EVIDENCE OF EVOLUTION
In order to become acquainted with the way the structures of animals provide evidences of evolution, it is by no means necessary to review the entire range of their forms, because research has discovered that the principles of relationship are universal among animals, and that any group of examples will demonstrate what is taught by comparative anatomy as a whole. The commonest creatures may serve us best in order that we may come to view evolution as a process that involves each and every living thing that we know, and not as something which belongs only to the remote and unknown past.
Let us begin with the common cat and the group of carnivora or flesh-eating animals to which it belongs. As we pass along the streets of the city, we will see many cats which differ in some details, though they resemble one another closely. While they vary somewhat in form, the range in this quality is not so noticeable as in the matter of color; some of them will be gray, some maltese, while others will be yellowish or black, and they will differ in the striped or spotted character of their coloration. We readily classify them all as "cats" in spite of their differences, because they are alike in so many ways that we have learned to associate as the distinguishing characteristics of these animals, and to label—"cat." The animals which we might see in a walk of several blocks may reasonably be regarded as offspring of the same pair of ancestors of a few years back, even though they are dissimilar. We all know that the kittens of one and the same litter vary: no two of them are ever exactly alike in color or disposition or voice or size, nor is any one identical with either of its parents, although it may be necessary to employ exact means of measuring them in order to demonstrate their variation. The fact of difference, then, is surely not inconsistent with even the closest ties of blood, and we do not need to go beyond the scope of daily observation to find that this is true in nature wherever we look.
Should we extend our observations so as to include the cats of Boston and Philadelphia and San Francisco, the animals would probably vary over a wider range, but they would be so similar to New York cats in their make-up that we would have no difficulty in regarding them and all the others of the United States as the descendants of a single pairs of ancestors, perhaps brought over in the "Mayflower." But why does this view seem justified? Because experience has taught us that the living things which resemble each other most closely are those which are most intimately bound by ties of blood and common heritage. It is "natural" for relatives to resemble one another more than persons not related, and for brothers and sisters to be more alike than cousins. Science does not refer to something outside everyday observation when it states that the possession by two animals of a great body of similar characters beneath their minor differences is an indication of their common ancestry.
Thus at the very outset our simple illustration establishes the most fundamental principle of comparative anatomy. Let us see how it works further. The Manx cat possesses an abbreviated tail, although in other respects it is practically the same as the familiar long-tailed form; the Angora and the Persian differ in having long hair. All of these animals are so much alike in so many respects, and so closely resemble the wild cats, that it is not unreasonable to regard them all as the descendants of the same original wild ancestors, and as the varying products of lines which branched out from the same stock in different directions and at different times. It is, in a word, their "cat-ness" which demonstrates their relationships. But common sense need not stop here. Guided by the facts of anatomical similarity, it convinces us that the dun-colored lion and puma, the striped tiger and the spotted leopard are simply cats of a larger growth whose remoter ancestry is one with that of the previously cited forms. Not until we explore and compare their several systems do we see how thoroughgoing is their uniformity in structural plan. And because reason justifies the view regarding the origin of domestic cats from wild ancestors, the evolution of all the various members of the cat tribe must be acknowledged. These animals exhibit a fundamental likeness, which, to employ a musical analogy, is the "theme" of "cat-ness," and they are so many variations of this theme.