From deposits found at Grimaldi, in the Grotte des Enfants and in the Grotte du Prince, it has long been said that men of early Mousterian times lived contemporary with the hippopotamus, the straight-tusked elephant, and Merck's rhinoceros in the genial climate of the Mediterranean Riviera. More recently the same animals have been found as far north as the Somme valley in the 'river-drifts' of Montières-les-Amiens.[(1)] Here, again, we find remains of the hippopotamus, the straight-tusked elephant, and its companion, Merck's rhinoceros, in Mousterian deposits, a surprising discovery, because it had always been supposed that a cold climatic period had set in all over western Europe even before the close of the Acheulean culture. But there is also evidence of a temperate climate still prevailing in the Thames valley in the period of the Mousterian 'floors.'[(2)] Again, along the Vézère valley, Dordogne, we find that at the station of La Micoque, where the industry marks the transition between late Acheulean and early Mousterian times, Merck's rhinoceros is found in the lowest layers associated with remains of the moose (Alces).

There is evidence that Merck's rhinoceros and the straight-tusked elephant lingered in western Europe during the whole period of the early development of the Mousterian industry. As observed above, these animals were hardier than the southern mammoth, which was the first of the Asiatic mammals to disappear, soon to be followed by its companion, the hippopotamus. Even after the advent of the closely associated tundra pair, the woolly mammoth and the woolly rhinoceros, Merck's rhinoceros persists, as, for example, in the deposits of Rixdorf, near Berlin, where this ancient type occurs in the same deposits with the woolly mammoth, the woolly rhinoceros, the reindeer, and the musk-ox, as well as with the forest forms, the moose, stag, wolf, and forest horse. The extreme northern latitude of this deposit explains the absence of the straight-tusked elephant, which may at the time have been living farther to the south. The same mingling of south and north Asiatic mammals is found at Steinheim, in the valley of the Murr, some degrees to the west and south of Rixdorf, not far from Göttingen, where we find Merck's rhinoceros[(3)] and the straight-tusked elephant in association with the woolly mammoth, the woolly rhinoceros, the giant deer, and the reindeer.

Thus the Neanderthal races were entering the Mousterian stage of culture during the close of the Third Interglacial Stage and during the early period of the advance of the ice-fields from the great centres in Scandinavia and the Alps. As these ice-fields slowly approached each other from the north and from the south a very great period of time must have elapsed during which all the south Asiatic mammals abandoned western Europe or became extinct, with the exception of the lions and hyænas, which became well fitted to the very severe climate that prevailed over Europe during the fourth glaciation, and even during the long Postglacial Stage which ensued. The large carnivora readily become thoroughly adapted to cold climates, as they subsist on animal life wherever it may be found; tigers of the same stock as those of India have been found as far north as the river Lena, in latitude 52° 25', where the climate is colder than that of Petrograd or of Stockholm, while the lion throve in the cold atmosphere of the upper Atlas range. Thus the cave-lion (Felis leo spelæa) and the cave-hyæna (H. crocuta spelæa) doubtless evolved an undercoating of fur as well as an overcoating of long hair, like the tundra mammals. In size the lion of this period in France often equalled and sometimes surpassed its existing relatives, the African and west Asiatic lion; it frequently figures in the art of the Upper Palæolithic artists and survived in western Europe to the very close of Upper Palæolithic times.

The Fourth Glaciation

Penck[(4)] has estimated that the first maximum of the fourth glaciation in the Alps was reached 40,000 years ago, and that after the recession period the second maximum ended not less than 20,000 years ago. This would extend the Mousterian industry over a very long period of time, for there can be no doubt that the Mousterian culture was practically contemporaneous with the fourth glaciation, even if a briefer period of time should be allotted to this great natural event.

The fourth glaciation, like the first, is believed to have been contemporaneous in Europe and North America,[(5)] a fact which is of especial importance to American anthropologists in connection with the question of the date of arrival of primitive man in America. In both countries the glaciation reached an early maximum, which was followed by a period of recession of the ice-fields, a time during which a somewhat more temperate climate prevailed, but this in turn gave way to a second advance of as great severity as the first.[AE]

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Fig. 94. Europe during the extension of the ice-fields and glaciers of the Fourth Glacial Stage. This is also supposed to have been a period of land depression and of extension of the inland seas of southern Europe. Britain was probably connected with France. The ice-covered areas in western Europe and Britain were far more limited than during the Third Glacial Stage, yet the climate appears to have been more severe than at any previous period. For the snow-level compare Fig. 13. Drawn by C. A. Reeds after Geikie and De Geer.

In the north, Scandinavia and Finland were again enshrouded in ice, and a great mer de glace occupied the basin of the Baltic Sea, sending its terminal moraines into Denmark and Schleswig-Holstein and over the northern provinces of Germany, but this great ice-field did not again become confluent with that of Great Britain.[(7)] At the commencement of the fourth glaciation large glaciers descended over the Scottish mountain valleys and filled many of them even to the sea; the coast subsided at least 130 feet in this region. In southern Britain along the valley of the Thames there spread an arctic flora, with the polar willow (Salix polaris) and the dwarf birch (Betula glandulosa); an arctic plant bed has also been discovered in the valley of the Lea. Thus the tundra climate extended from the Scottish lowlands to the south of England, the land being bleak and almost treeless.[(8)] This, we believe, was also the period of the arctic flora at Hoxne, Suffolk, and of the arctic plant bed in the valley of the Thames. At this time the valley was frequented by the reindeer, the woolly rhinoceros, and the mammoth, whose remains are entombed in the low-level alluvia swept down from the sides of the valley, so that the remains of this arctic fauna may in places actually overlie those of the more deeply buried and far more ancient warm Asiatic fauna of Chellean times. Like the Somme, the Thames[(9)] was then from 10 to 25 feet below its present level, the bottom having since silted up with alluvial soil.