The grinding-teeth do not all show this massive size and columnar form, for about fifty per cent of the Krapina teeth have distinct roots and are more like normal modern grinders. In the Neanderthaloids of Spy the teeth are small and the roots are of moderate size.[(55)]
This study of the forehead and of the eyebrow ridges, of the great depth of the face, and of the peculiarly high, square form of the eye sockets prepares us for a profile view of the skull of La Chapelle in contrast with that of the most highly developed and intellectual European type, namely, the profile of the distinguished American palæontologist, the late Professor Edward D. Cope, who bequeathed his skull and skeleton for purposes of scientific study and comparison. In La Chapelle we at once notice the platycephaly, or flattening of the skullcap, the retreating forehead, the great prominence of the eyebrow ridges resembling that of the anthropoid apes, the lengthening of the face as compared with the flattening of the cranium, the great prominence or prognathism of the face as a whole, and the special prominence of the rows of cutting teeth as compared with the vertical or indrawn line, and the recession of the tooth row in the Cope profile. This comparison also brings out the striking contrast between the high chin prominence of Homo sapiens and the deeply receding chin of the Neanderthals. The contrast is hardly less remarkable in the superior view of the skull in which the Neanderthal type is seen to be extremely dolichocephalic, the back of the skull being relatively broad and the front narrowing in the region of the forebrain until it suddenly expands in the prominent supraorbital processes.
Fig. 115. Skull of La Chapelle-aux-Saints (outline) in comparison with that of a high modern type (shaded); illustrating the projecting eyebrows and prognathous, ape-like face of the Neanderthaloids. After Boule. One-quarter life size.
As shown in the diagram on page 8, Fig. 1, the greatest length of the Neanderthal skull is found on the horizontal line directly through the brain chamber, known as the glabella-inion line, a line drawn from a prominence between the eyebrow ridges to a point at the back of the skull known as the external occipital protuberance, or inion. This is also the longest line in the skulls of Spy and of La Chapelle, as well as of the anthropoid apes,[(56)] but in the north Australian skull, Fig. 1, owing to the greater expansion of the upper part of the brain, the greatest length of the skull is at a point considerably above the glabella-inion line. The median section of the skull of the chimpanzee, of the Neanderthal, and of the north Australian displays in a very striking manner the generalization made by Schwalbe, in 1901, that the Neanderthal skull is truly an intermediate or half-way form between that of the anthropoid apes and that of Homo sapiens. We observe in this illuminating section the growth of the dome of the skull, that is, the great brain-bearing cavity above the glabella-inion line g-i, by noting the contrast in the length of the vertical line of the cranial height, as compared with the space below the glabella-inion line indicated by the letters. This very important vertical line terminates below at the opening, where the spinal cord enters the base of the brain (see Fig. 1).
Fig. 116. Top view of three skulls—of a chimpanzee (left), of the man of La Chapelle-aux-Saints (centre), and of a modern Frenchman (right)—showing the retreat of the projecting face and prominent eyebrow ridges. After Boule.
In many characteristics the Neanderthal skull is shown to be nearer to that of the anthropoid apes than to that of Homo sapiens. This conclusion arrived at by Schwalbe, in 1901,[(57)] has been more than confirmed by Boule's masterly study[(58)] of the very complete skull of La Chapelle. After his detailed review, he concludes: As to the unity of the Neanderthal head form, these features are not peculiar to the skull of La Chapelle; in every case they are also found in the skulls of Neanderthal, Gibraltar, Spy, Krapina, La Ferrassie, which witness to the homogeneity of that human fossil type called Neanderthal. These features show a structural affinity between the fossil men of the Mousterian period and the anthropoid apes. It must be noted that many of these features may be found also in recent human skulls of the inferior races, but that they are very rare, very scattered, very isolated, and occur only as aberrations. It is the accumulation of all these features in every skull of a whole series which constitutes an assemblage entirely new and of great importance. In the skull, as in other parts of the anatomy of the Neanderthals, we should not expect to find every character intermediate between the anthropoids and recent man. The long Neanderthal face is somewhat similar to that of the Eskimo and is in contrast with the very short face of the existing Australians and Tasmanians. The depression at the root of the nose, just below the glabella, is very marked in all Neanderthals; there is less of the nose bridge than in any recent races, except those of the male Australians, yet the nose is not flattened but somewhat arched or aquiline. This feature is not characteristic of all the anthropoid apes, and in this respect the Neanderthals, Australians, and Tasmanians are more different from the anthropoid apes than are some of the white races; thus the Neanderthal nose, far from resembling that of the anthropoids, differs from it more than does that of some recent human types.[(59)] Many anatomists, following Huxley, have described the Australian and Tasmanian skulls as more or less Neanderthaloid, and some authors have gone so far as to regard these races as surviving Neanderthals. It is true that some of the skulls in these existing races are extraordinarily platycephalic and show a retreating forehead, that others show supraorbital ridges almost as prominent as in the Neanderthals, that sometimes the prominence of the occipital inion is very marked, that certain jaws show a very retreating chin. Thus one or another of these Neanderthal features has been observed in these lower existing races, but all of these characteristics have never been combined in one race as constant features, and invariably associated, as in all the skulls of the Neanderthals known to us.
Fig. 117. Scale of ascent indicated in the skull form of eleven races of fossil and living men, based on the result of twelve different characters of comparison. At the bottom stands the anthropoid ape, and above this Pithecanthropus, the ape-man of Java. A wide range is observed between the Neanderthaloid skulls of Gibraltar and of Spy-Neanderthal. Not far above these in the scale of ascent stand the modern Australians and the recently extinct Tasmanians. Above these low races are found the fossil Upper Palæolithic races of Galley Hill, Brüx, Brünn, and Předmost. At the top stand the modern European races, beside which the Upper Palæolithic Crô-Magnon race takes a high rank. After Büchner.