In the 1950-51 season, small young of the summer brood seemed unusually numerous. In the 1951-52 period, young of both age classes were relatively scarce and old adults made up an unusually high proportion of the population. Excluding the 14 marked pouch young that were later recaptured, there were only four of the total of 106 that were trapped in each of two seasons. One young less than a quarter grown, that was accidentally caught in a live-trap set for woodrats, was recaptured as a breeding adult the following winter. An adult male and two adult females each caught in the 1949-50 season were each recaptured repeatedly in the 1950-51 season. Ninety-five per cent replacement of the breeding population by the following breeding season is indicated by our figures. Only 3 (or 5 per cent) of the individuals of the population trapped and marked in the season of 1949-50, were recaptured among the 62 opossums recorded in the two subsequent seasons. Various mortality factors including predation, disease, and accidents account for some 70 per cent. These are replaced by first-year young which make up the greater part of the breeding population. The remaining 25 per cent presumably shift their ranges sufficiently in the course of a year to have moved beyond the limits of an area of the size encompassed by the present study.

POPULATION DENSITY

No precise measurement of the population density on the study area was obtained. It was not practical to capture every individual present there, and rapid population turnover, due to mortality and wandering, obscured the trends. The information obtained concerning movements of opossums suggest that one may habitually forage as much as 900 feet from its home base. Assuming that 900 feet is the typical cruising radius, the areas drawn upon by the trap lines in the three different seasons were approximately as follows: 1949-50—400 acres; 1950-51—350 acres; 1951-52—220 acres. In these same three seasons the numbers of opossums caught were, respectively, 46, 37, and 30. If these figures represent the numbers actually present, densities of one to 8.7 acres, one to 9.5 acres, and one to 7.3 acres are indicated. However, some opossums using the area probably were missed; and on the other hand, not all those caught in the course of a season were present there simultaneously. Many of those present early in the season would have moved away a few months later, and others would have moved in, replacing them. The number present at any one time could scarcely have been more than half the number caught in the entire season.

Census With Half-monthly Sampling Periods

Sampling periodNumber of
individuals taken
in period
Number of
individuals taken
in following period
Number of
recatures in
following period
Computed population
for sampling period
Early November 194937121
Late November 194978318.7
Early December 1949811329.3
Late December 1949117419.2
Early January 195073121
Early March 195058220
Late March 195086316
Early April 195063118
Late April 19503629
Early May 19506329
Early November 19501313
Late December 195036118
Early February 1951413317.3
Late February 1951136326
Early March 19516438
Late March 195145210
Early April 19515115
Late April 19511515
Early May 19515327.5
Early February 195294218
Late February 195249136
Early March 195296227
Late March 195265215

Census with Monthly Sampling Periods

Sampling periodNumber of
individuals taken
in period
Number of
individuals taken
in following period
Number of
recatures in
following period
Computed population
for sampling period
November 1949916721
December 1949169348
March 1950119333
April 195097232
October 19509339
November 19503319
December 19503737
January 1951714333
February 1951147425
March 195175312
April 195156310
November 195136118
December 195165130
January 1952511318
February 19521113436
March 1952139523
April 195293127

Crude census-figures were obtained by utilizing the Lincoln Index and computing the total on the basis of the ratio of marked (and recognizable) individuals to others caught in a sampling period. A large number of census figures were obtained over the three-year period of the study. Each separate census, however, was based on an inadequate sample as the number of marked individuals taken at each sampling, as recaptures from the previous sampling period, varied from one to five. While little confidence can be placed in any one census computation, the trends of figures from series of such computations reveal the approximate number of opossums on the area if due allowance is made for certain distorting factors. Presumably the differences in figures obtained at different samplings result chiefly from the margin of error in the data, although it is true that there is rapid change in the actual number of opossums.

The number of active opossums in the region of the study reaches a peak in late summer and early fall, when second litters of young have grown large enough to become independent. At this season the population contains a high proportion of young of the year. During the ensuing months of fall and winter there is a steady decrease in numbers, through various mortality factors, with no replacement until young are born about the first week of March. These young do not become independent until late May or early June, and during the intervening months there is a further reduction of the adults and yearlings, so that the active population reaches its annual low point in late spring. At that time of year most opossums are in poor physical condition.