Assuming, however, that the facilities of immigration had become adequate; which would be the first mammals to arrive and live? Not large herbivores; for they would be soon drowned if by any accident carried out to sea. Not the carnivora; for these would lack appropriate food, even if they outlived the voyage. Small quadrupeds frequenting trees, and feeding on insects, would be those most likely both to be drifted away from their native lands and to find fit food in a new one. Insectivorous mammals, like in size to those found in the Trias and the Stonesfield slate, might naturally be looked for as the pioneers of the higher vertebrata. And if we suppose the facilities of communication to be again increased, either by a further shallowing of the intervening sea and a consequent multiplication of islands, or by an actual junction of the new continent with an old one, through continued upheavals; we should finally have an influx of the larger and more perfect mammals.

Now rude as is this sketch of a process that would be extremely elaborate and involved, and open as some of its propositions are to criticisms which there is no space here to meet; no one will deny that it represents something like the biologic history of the supposed new continent. Details apart, it is manifest that simple organisms, able to flourish under simple conditions of life, would be the first successful immigrants; and that more complex organisms, needing for their existence the fulfilment of more complex conditions, would afterwards establish themselves in something like an ascending succession. At the one extreme we see every facility. The new individuals can be conveyed in the shape of minute germs; these are infinite in their numbers; they are diffused in the sea; they are perpetually being carried in all directions to great distances by ocean-currents; they can survive such long journeys unharmed; they can find nutriment wherever they arrive; and the resulting organisms can multiply asexually with great rapidity.

At the other extreme, we see every difficulty. The new individuals must be conveyed in their adult forms; their numbers are, in comparison, utterly insignificant; they live on land, and are very unlikely to be carried out to sea; when so carried, the chances are immense against their escape from drowning, starvation, or death by cold; if they survive the transit, they must have a pre-existing Flora or Fauna to supply their special food; they require, also, the fulfilment of various other physical conditions; and unless at least two individuals of different sexes are safely landed, the race cannot be established. Manifestly, then, the immigration of each successively higher order of organisms, having, from one or other additional condition to be fulfilled, an enormously-increased probability against it, would naturally be separated from the immigration of a lower order by some period like a geologic epoch.

And thus the successive sedimentary deposits formed while this new continent was undergoing gradual elevation, would seem to furnish clear evidence of a general progress in the forms of life. That lands thus raised up in the midst of a wide ocean, would first give origin to unfossiliferous strata; next, to strata containing only the lowest marine forms; next, to strata containing higher marine forms, ascending finally to fish; and that the strata above these would contain reptiles, then small mammals, then great mammals; seems to us to be demonstrable from the known laws of organic life.

And if the succession of fossils presented by the strata of this supposed new continent, would thus simulate the succession presented by our own sedimentary series; must we not say that our own sedimentary series very possibly records nothing more than the phenomena accompanying one of these great upheavals? We think this must be considered not only possible, but highly probable: harmonizing as it does with the unavoidable conclusion before pointed out, that geological changes must have been going on for a period immeasurably greater than that of which we have records. And if the probability of this conclusion be admitted, it must be admitted that the facts of Palæontology can never suffice either to prove or disprove the Development Hypothesis; but that the most they can do is, to show whether the last few pages of the Earth's biologic history are or are not in harmony with this hypothesis—whether the existing Flora and Fauna can or can not be affiliated upon the Flora and Fauna of the most recent geologic times.

[T] Sir Charles Lyell is no longer to be classed among Uniformitarians. With rare and admirable candour he has, since this was written, yielded to the arguments of Mr. Darwin.

IX.
THE DEVELOPMENT HYPOTHESIS.

In a debate upon the development hypothesis, lately narrated to me by a friend, one of the disputants was described as arguing, that as, in all our experience, we know no such phenomenon as transmutation of species, it is unphilosophical to assume that transmutation of species ever takes place. Had I been present, I think that, passing over his assertion, which is open to criticism, I should have replied that, as in all our experience we have never known a species created, it was, by his own showing, unphilosophical to assume that any species ever had been created.

Those who cavalierly reject the Theory of Evolution, as not adequately supported by facts, seem quite to forget that their own theory is supported by no facts at all. Like the majority of men who are born to a given belief, they demand the most rigorous proof of any adverse belief, but assume that their own needs none. Here we find, scattered over the globe, vegetable and animal organisms numbering, of the one kind (according to Humboldt), some 320,000 species, and of the other, some 2,000,000 species (see Carpenter); and if to these we add the numbers of animal and vegetable species that have become extinct, we may safely estimate the number of species that have existed, and are existing, on the Earth, at not less than ten millions. Well, which is the most rational theory about these ten millions of species? Is it most likely that there have been ten millions of special creations? or is it most likely that by continual modifications, due to change of circumstances, ten millions of varieties have been produced, as varieties are being produced still?