Dwelling for a moment on this last illustration of the class of difficulties described, let us ask how we are to interpret the development of the musical faculty. I will not enlarge on the family antecedents of the great composers. I will merely suggest the inquiry whether the greater powers possessed by Beethoven and Mozart, by Weber and Rossini, than by their fathers, were not due in larger measure to the inherited effects of daily exercise of the musical faculty by their fathers, than to inheritance, with increase, of spontaneous variations; and whether the diffused musical powers of the Bach clan, culminating in those of Johann Sebastian, did not result in part from constant practice; but I will raise the more general question—How came there that endowment of musical faculty which characterizes modern Europeans at large, as compared with their remote ancestors. The monotonous chants of low savages cannot be said to show any melodic inspiration; and it is not evident that an individual savage who had a little more musical perception than the rest, would derive any such advantage in the maintenance of life as would secure the spread of his superiority by inheritance of the variation. And then what are we to say of harmony? We cannot suppose that the appreciation of this, which is relatively modern, can have arisen by descent from the men in whom successive variations increased the appreciation of it—the composers and musical performers; for on the whole, these have been men whose worldly prosperity was not such as enabled them to rear many children inheriting their special traits. Even if we count the illegitimate ones, the survivors of these added to the survivors of the legitimate ones, can hardly be held to have yielded more than average numbers of descendants; and those who inherited their special traits have not often been thereby so aided in the struggle for existence as to further the spread of such traits. Rather the tendency seems to have been the reverse.
Since the above passage was written, I have found in the second volume of Animals and Plants under Domestication, a remark made by Mr. Darwin, practically implying that among creatures which depend for their lives on the efficiency of numerous powers, the increase of any one by the natural selection of a variation is necessarily difficult. Here it is.
“Finally, as indefinite and almost illimitable variability is the usual result of domestication and cultivation, with the same part or organ varying in different individuals in different or even in directly opposite ways; and as the same variation, if strongly pronounced, usually recurs only after long intervals of time, any particular variation would generally be lost by crossing, reversion, and the accidental destruction of the varying individuals, unless carefully preserved by man.”—Vol. ii, 292.
Remembering that mankind, subject as they are to this domestication and cultivation, are not, like domesticated animals, under an agency which picks out and preserves particular variations; it results that there must usually be among them, under the influence of natural selection alone, a continual disappearance of any useful variations of particular faculties which may arise. Only in cases of variations which are specially preservative, as for example, great cunning during a relatively barbarous state, can we expect increase from natural selection alone. We cannot suppose that minor traits, exemplified among others by the æsthetic perceptions, can have been evolved by natural selection. But if there is inheritance of functionally-produced modifications of structure, evolution of such minor traits is no longer inexplicable.
Two remarks made by Mr. Darwin have implications from which the same general conclusion must, I think, be drawn. Speaking of the variability of animals and plants under domestication, he says:—
“Changes of any kind in the conditions of life, even extremely slight changes, often suffice to cause variability.... Animals and plants continue to be variable for an immense period after their first domestication; ... In the course of time they can be habituated to certain changes, so as to become less variable; ... There is good evidence that the power of changed conditions accumulates; so that two, three, or more generations must be exposed to new conditions before any effect is visible.... Some variations are induced by the direct action of the surrounding conditions on the whole organization, or on certain parts alone, and other variations are induced indirectly through the reproductive system being affected in the same manner as is so common with organic beings when removed from their natural conditions.”—(Animals and Plants under Domestication, vol. ii, 270.)
There are to be recognized two modes of this effect produced by changed conditions on the reproductive system, and consequently on offspring. Simple arrest of development is one. But beyond the variations of offspring arising from imperfectly developed reproductive systems in parents—variations which must be ordinarily in the nature of imperfections—there are others due to a changed balance of functions caused by changed conditions. The fact noted by Mr. Darwin in the above passage, “that the power of changed conditions accumulates; so that two, three, or more generations must be exposed to new conditions before any effect is visible,” implies that during these generations there is going on some change of constitution consequent on the changed proportions and relations of the functions. I will not dwell on the implication, which seems tolerably clear, that this change must consist of such modifications of organs as adapt them to their changed functions; and that if the influence of changed conditions “accumulates,” it must be through the inheritance of such modifications. Nor will I press the question—What is the nature of the effect registered in the reproductive elements, and which is subsequently manifested by variations?—Is it an effect entirely irrelevant to the new requirements of the variety?—Or is it an effect which makes the variety less fit for the new requirements?—Or is it an effect which makes it more fit for the new requirements? But not pressing these questions, it suffices to point out the necessary implication that changed functions of organs do, in some way or other, register themselves in changed proclivities of the reproductive elements. In face of these facts it cannot be denied that the modified action of a part produces an inheritable effect—be the nature of that effect what it may.
The second of the remarks above adverted to as made by Mr. Darwin, is contained in his sections dealing with correlated variations. In the Origin of Species, p. 114, he says—
“The whole organization is so tied together during its growth and development, that when slight variations in any one part occur, and are accumulated through natural selection, other parts become modified.”