Fig. 253.
Composite flowers furnish evidence so nearly allied to that which clustered flowers furnish, that we may fitly glance at them under the same head. Such a common type of this order as the Sun-flower, exemplifies the extremely marked difference which arises in many of these plants between the closely-packed internal florets, each similarly circumstanced on all sides, and the external florets, not similarly circumstanced on all sides. In Fig. [253], representing the inner and outer florets of a Daisy, the contrast is marked between the small radial corolla of the one and the larger bilateral corolla of the other. In many cases, however, this contrast is less marked: the inner florets also having their outward-growing prolongations—a difference possibly related to some difference in the habits of the insects that fertilize them. Nevertheless, these composite flowers which have inner florets with strap-shaped corollas outwardly directed, equally conform to the general principle; both in the radial arrangement of the assemblage of florets, and in the bilateral shape of each floret; which has its parts alike on the two sides of a line passing from the centre of the assemblage to the circumference. Certain other members of this order fulfil the law somewhat differently. In Centaurea, for instance, the inner florets are small and vertical in direction, while the outer florets are large and lateral in direction. And here may be remarked, in passing, a clear indication of the effect which great flexibility of the petals has in preventing a flower from losing its original radiate form; for while in C. cyanus, the large outward-growing florets, having short, stiff divisions of the corolla, are decidedly bilateral, in C. scabiosa, where the divisions of the corolla are long and flexible, the radial form is scarcely at all modified. On bearing in mind the probable relations of the forms to insect-agency, the meaning of this difference will not be difficult to understand.[38]
§ 236. In extremely-varied ways there are thus re-illustrated among flowers, the general laws of form which leaves and branches and entire plants disclose to us. Composed as each cluster of flowers is of individuals that are originally similar; and composed as each flower is of homologous foliar organs; we see both that the like flowers become unlike and the like parts of each flower become unlike, where the positions involve unlike incidence of forces. The symmetry remains radial where the conditions are equal all round; shows deviation towards two-sidedness where there is slight two-sidedness of conditions; becomes decidedly bilateral where the conditions are decidedly bilateral; and passes into an unsymmetrical form where the relations to the environment are unsymmetrical.
CHAPTER XI.
THE SHAPES OF VEGETAL CELLS.
§ 237. We come now to aggregates of the lowest order. Already something has been said ([§ 217]) concerning the forms of those morphological units which exist as independent plants. But it is here requisite briefly to note the modifications undergone by them where they become components of larger plants.
Fig. 254.
Of the numerous cell-forms which are found in the tissues of the higher plants, it will suffice to give, in Fig. [254], representing a section of a leaf, a single example. In this it will be seen that the cells forming the upper and lower surfaces, a and b, have differences of shape related to differences in the incidence of forces: they are more or less flattened in relation to the environment. The underneath cells at c, form a class which, similarly exposed to light at their outer ends, and, as we may assume, largely developed in adjustment to their active assimilative functions, are, by mutual pressure, made to grow more in the direction of their lengths than in the direction of their breadths. Then on the other side we see that the cells d, next above the outer layer, while approximately similar, become more and more dissimilar as they diverge from the surface, and are quite irregular in the interior e, where there is no definiteness in the conditions to which they are exposed. Thus the divergences of these cells from primordial sphericity are such as correspond with unlikenesses in their circumstances. And throughout the more complex modifications which the cells of other tissues exhibit, the like correspondences hold.
Figs. 32–35.