Fig. 2.

Palæontological breaks or breaks in the succession of organisms are in many cases, the result of physical breaks, and accordingly it is often possible to separate one set of strata from another by the existence of a combined physical and palæontological break between them. It is by no means necessary however that a physical break should be accompanied by a break in succession of the organisms, and the latter may also occur without the former. It was once maintained that a palæontological break was due to the complete and sudden extinction of a fauna and its entire replacement by a new one, but this is far from true, and accordingly the breaks differ in degree. Study of the strata shows that when the succession is not to any extent interrupted, the species do not appear simultaneously, but come in at different horizons, and they disappear in the same way. In [Figure 2] let A represent a set of conformable strata ab ... k, and suppose the vertical lines represent the ranges of the various species found in these strata. It will be seen that of 27 species whose range is shown only 2 pass through the whole thickness, so that the fauna of k is very different from the fauna of a, nevertheless the fauna of each stratum is closely similar to that of the underlying as well as to that of the overlying stratum, and though most of the species of k are different from those of a, this need not be the case with the genera. The fauna of the set of strata would contain every species whose range is represented, and for convenience' sake it might be said to be composed of sub-faunas, one of which occurs in each division ab ..., but the separation into sub-faunas would be artificial and merely for convenience' sake, for there is no break between any two sub-faunas. Turning now to B ([Fig. 2]), an attempt is made there to show what happens when there has been a physical break, resulting in the denudation of the strata ghik, and the deposition of another set op ... unconformably upon those deposits of the earlier set which have not been denuded. As the result of this we note, first, that the relics of organisms which existed in the area during the deposition of ghik, and were entombed in those strata, are destroyed by the processes of denudation, and a large number of organisms which lived long after the deposition of f, and disappeared not simultaneously but at different times during the period when denudation was in operation, seem to become extinct simultaneously at the top of f, though, if we could visit an area which was receiving sediment during the period of denudation, we should find them dying out in the rocks of that region at different levels. Furthermore, whilst denudation is going on, a longer or shorter period of time elapses, during which the upheaved area receives no deposit, and accordingly no organisms which lived during that period are preserved in the upheaved area. During this time a set of deposits lmn may have been laid down elsewhere, and besides the gradual disappearance of some of the organisms of ab ... k, there will have been a gradual appearance of new species. When the upheaved area is once more submerged, a new set of deposits op ... is accumulated in it, and the species which gradually appeared in adjoining regions will now migrate to it, and will seem to come in simultaneously at the bottom of o; accordingly we may find that there is not a single species which passes through from f to o and the palæontological break in this area is complete, though it is clear that it only implies local change, and that we may and indeed must find intermediate forms in other regions which fill up the gap.

As an illustration of the local character of a palæontological break we may cite the case of the Carboniferous and Permian systems of Britain. These rocks are separated from one another in our area by a physical and palæontological break, but in parts of India, and other places, we find a group of rocks now known as the Permo-Carboniferous rocks which contain a fauna intermediate in character between those of the Permian and Carboniferous systems, and a study of this fauna shows that the hiatus which exists locally is filled by the species contained in the Permo-Carboniferous rocks.

A palæontological break may, like a physical one, result from depression of the ocean-floor to so great a depth, that no organisms are preserved there during the period of great depression, and the remarks made concerning a depression of this nature when speaking of physical breaks will apply here also.

A local palæontological break may result owing to physical changes without the production of an unconformity in the area, or its submergence to a great depth, or if an unconformity is found, the break may be more marked owing to other physical changes. The difference between the Upper and Lower Carboniferous faunas is very marked in England, where the Upper Carboniferous beds were deposited under physical conditions different from those of the Lower Carboniferous, and accordingly the corals, crinoids and other open-water animals which flourished in Lower Carboniferous times are rare or altogether absent in the higher rocks. Where the change of conditions did not occur to a great extent as in parts of Spain and North America, the similarity between the two faunas is much more pronounced. Again, there is an unconformity between the Cretaceous and Eocene beds of England, which is accompanied by a palæontological break, but this break is more pronounced owing to difference of physical conditions, for we find abundance of gastropods in the lower Tertiary beds, and a rarity of these shells at the top of the chalk of England, though where physical conditions were favourable for the growth of gastropods, their shells are found in the higher strata of chalk age, and the palæontological break is not so apparent.

A palæontological break may occur also as the result of climatic change, though actual instances of this occurrence are much more difficult to detect owing to the general absence of any evidence of climatic change other than that supplied by the organisms themselves. Still, when no physical break exists, and the lithological characters of a group of sediments remain constant throughout, indicating the prevalence of similar physical conditions through the period of deposition of the sediments, if the fauna suddenly changes, there must have been cause for the change, and in the absence of any other cause which is likely to produce the change, alteration of the character of the climate may be suspected.

It follows from the observations which have been made, that although the rocks of the Geological Column may be divided into systems owing to the existence of physical and palæontological breaks, and this classification may be and has been applied generally, the line of demarcation between the rocks of two systems will be a purely conventional one, where there is no break, and, to avoid confusion, that line when once drawn should be adopted by everyone, unless good cause can be shown for its abandonment.