The characters of the Ordovician trilobites have already been noticed. These organisms are abundant, and occur in sediments of all kinds. Of other groups, the significance of the radiolaria has been referred to above. Corals occasionally form reef-like masses of limestone as in the limestones of the Caradoc epoch; the echinoderms are well represented, cystids being locally abundant; of the crustacea, many remains of tests of phyllocarida have been recorded; the brachiopods are very abundant, and of the mollusca, lamellibranchs, gastropods and cephalopods all occur with frequency though none of these groups is very prevalent. Certain forms have been referred to pteropods though with doubt, and other shells seem to be referable to the heteropods. The existence of vertebrates during Ordovician times is not, in the opinion of many geologists, proved, though remains of fishes have been recorded from the Ordovician strata of North America; but it is desirable that more evidence of this occurrence should be given[79].

[79] Walcott, C. D., "Preliminary Notes on the Discovery of a Vertebrate Fauna in Silurian (Ordovician) Strata," Bulletin Geol. Soc. America, vol. III. p. 153.

The distribution of the Ordovician faunas like that of the sediments points to the prevalence of open ocean conditions over wide areas during the period, with occasional approaches to land, which was often of a volcanic nature. Around this land clustered the ordinary invertebrates, building up coral-reefs and shell-banks, whilst away in the open oceans the graptolites floated, almost alone, and sank to the ocean floor after death.


[CHAPTER XVI.]

THE SILURIAN SYSTEM AND THE CHANGES WHICH OCCURRED IN BRITAIN AT THE CLOSE OF SILURIAN TIMES.

Classification. The Silurian system was originally divided by its founder, Sir R. I. Murchison, into three series, as follows:—

LudlowSeries
Wenlock"
Llandovery"

The term May Hill, proposed by Sedgwick, is sometimes used as synonymous with Llandovery. This classification omits a somewhat important set of beds intercalated between those of the Llandovery and Wenlock series known as the Tarannon shales, and in Britain if we were to classify afresh, it would be more convenient to include some of the beds formerly referred to the Ludlow in the Wenlock. I shall, however, adopt the old and well-established classification, adding the term Tarannon to Llandovery, and speaking of the Llandovery-Tarannon series. The nature of the two classifications is shown in the following table: