XX.—ACTION OF LIGHT AND WARMTH ON AUTONOMOUS ACTIVITY

By

Sir J. C. Bose.

In the preceding Paper I have shown the essential similarity of effect of stimulus on autonomous activity of the Desmodium leaflet, and of the growing organ. It was shown how stimulus revived the pulsatory activity of Desmodium leaflet in a state of standstill, in the same way as it renewed the arrested growth-activity.

THE OSCILLATING RECORDER.

The in­ves­ti­ga­tion of this subject was rendered possible by the successful device of my Oscillating Recorder. A very light glass fibre was used for the construction of the lever, which was supported on jewel bearings. The short arm of the lever was 2 cm. in length, and the long arm 8 cm. This gave a mag­ni­fi­ca­tion of 4 times. But it is quite easy to increase the mag­ni­fi­ca­tion to 10 times or more.

The pull exerted by the pulsating leaflet is extremely slight, and the relatively heavy lever made of steel wire used in the Resonant Recorder is not well-suited for our purpose. The pulsation of the leaflet is relatively slow, being once in two minutes or so. The intermittent contact of ten times in a second, given by the Resonant Recorder, is therefore too quick. In the Oscillating Recorder the intermittence was, therefore, reduced to once in a second, or once in five seconds, the recording plate itself being made to move to-and-fro at this rate. The carrier of the plate-holder slides backwards and forwards on ball bearings; a wheel in the clockwork connected with an eccentric is released periodically, at intervals which may be varied between one and five seconds. By the action of the eccentric, the plate carrier approaches the writing lever with diminish­ing speed till the movement is zero at the contact. This contrivance is essential, since any sudden shock of the plate against the lever is apt to give rise to after-vibrations of the writer. The plate carrier is quickly withdrawn after the production of a dot on the smoked glass plate by contact with the writing lever.

The clockwork is governed by a revolving fan which can be gradually opened out by a regulating screw. The speed can thus be adjusted within wide limits, and maintained constant and at any desired speed. A second set of wheels connected with the clockwork moves the plate-holder in a lateral direction. A series of records may thus be taken for fifteen minutes, half an hour, or an hour.

The record obtained in this way is very perfect. Not only is the effect of an external agent shown by variation in the amplitude and frequency of pulsations, but the change of speed in any phase of the pulse becomes automatically recorded.