Fig. 10.—Skeleton of Cassowary, showing reduced wing-bones (a piece of black paper is placed under them).

From Dendy's Outlines of Evolutionary Biology (Constable).

Another and a very strong evidence of Evolution is to be found in what are termed vestigeal structures, two of which are illustrated in Figs. 9 and 10. They are, for the most part, obviously useless, and their occurrence has never been satisfactorily explained except by supposing them to be remnants of organs that were functional in the past history of their possessors' race. The appendix of man, for instance, is not only useless, but is frequently a source of danger. But its presence is readily explained by supposing that it represents the blind-gut, which is large and functional in many of the lower animals. Again, how should we account for the presence of small functionless wing-bones in the cassowary, unless by supposing that its ancestors were accustomed to fly like ordinary birds? How should we explain the bones which represent the hind limbs of the whale, unless by regarding the whale as descended from an animal which had functional hind limbs, or the representatives of eyes in animals that live in the dark, unless by supposing that these are descended from ancestors which saw? It has been well said that the bodies of many animals are veritable antiquarian museums, filled with relics of their own ancestors.

The next argument for Evolution to which we would refer is based on the similar structure and origin of organs or members that have entirely different uses. In Fig. 11 are figured the bones of the fore limbs of four different mammals, a whale, a bat, a dog, and man. The first is used for swimming, the second for flight, the third for locomotion on land, and the fourth as a grasping and holding organ. If these organs had been specially designed, each for its specific purpose, we should expect to find fundamental differences in structure. Actually the general arrangement of bones is the same in each case. A fact like this points strongly to a common origin of the four types mentioned, and to a general primitive arrangement of the bones of the limb. This primary type, it seems natural to suppose, has been modified for various special purposes in many different directions, the general features remaining recognisable. Many other cases of homology, or similarity of structure and origin, in organs whose function is dissimilar, might be quoted. Thus the poison gland of the poison snakes is not an organ which has been specially developed, but is a modified portion of one of the salivary glands. The hoof of the horse and the finger nail of man can evidently be satisfactorily explained as modifications of a general type of terminal claw, and the scales of the scaly ant-eater and the quills of the porcupine are only modified hairs. The significance of facts like these, when carefully considered, is very great.

Fig. 11.—The bones of the fore limbs of (a) whale, (b) bat, (c) dog, and (d) man, showing essential similarity in arrangement.

Fig. 12.—Distribution of Marsupials or pouch-bearing animals.

Australia, New Guinea, etc. 36 Genera. 144 Species.
America 3 Genera. 28 Species.

The study of the geographical distribution of animals has brought forth a great mass of facts which, considered by themselves, seem chaotic and meaningless, but which, in the light of Evolution, are full of significance. Observe, for example, the distribution of the Marsupials or pouch-bearing animals, shown on the accompanying map (Fig. 12). Australia is full of them, while they are relatively meagrely represented in a few other parts of the world. At the same time the greater and higher group of mammals was represented in Australia, at the time of its discovery, only by the bushman and his dog and a few species of mice. It is not as if the Australian environment were specially well adapted for marsupials, or specially ill-adapted for higher mammals; for the sheep has proved itself splendidly adapted for the conditions, and the rabbit most inconveniently so. Why, then, this curious state of affairs? It is an undoubted fact that the marsupials are both lower in their position in the animal kingdom, and older, than the main group to which all our European mammals belong. Now it is believed that Australia was once connected by land with the Asiatic Continent, and that it was finally separated from it before the higher mammals were in existence. The great step of further progress occurred elsewhere than in Australia, and the mammals of the latter continent were left in their obsolete condition, preserved through lack of competition of that higher type which elsewhere became dominant.