3. It has been found that young animals, birds, etc., depend upon the example and instruction of adults for the first performance of many actions that seem to be instinctive. This dependence may exist even in cases in which there is yet a congenital tendency to perform the action. Many birds, for example, have a general instinct to build a nest; but in many cases, if put in artificial circumstances, they build imperfect nests. Birds also have an instinct to make vocal calls; but if kept from birth out of hearing of the peculiar notes of their species, they come to make cries of a different sort, or learn to make the notes of some other species with which they are thrown.

4. The principal agency for the learning of the animals, and for the supplementing of their instincts, is Imitation. The sight of certain movements on the part of the adult animals, or the hearing of their cries, calls, notes, etc., leads the young to fall into an imitation of these movements or vocal performances. The endowment which such a young animal has in the direction of making movements and cries similar to those of his species aids him, of course, in imitating these in preference to others. So the endowment and the tendency to imitate directly aid each other in all such functions, and hurry the little creature on in his acquisition of the habits of his species. We find young animals clinging even in their imitations pretty closely to their own proper fathers and mothers, who are thus enabled to bring them up comme il faut.

5. There is every reason to think, moreover, that the tendency to imitate is itself instinctive. Young animals, notably the monkey and the child, fall spontaneously to imitating when they reach a certain age. Imitation shows itself to be instinctive in the case of the mocking bird, the parrot, etc. Furthermore, the mechanism of this function of imitation is now very well known. The principle of psychology recognised above under the phrase Kinæsthetic Equivalents, teaches us that the idea of a movement, coming into the mind through sight or some other sense, stirs up the proper apparatus to bring about the same movement in the observer. This we see in the common tendency of an audience to repeat the gestures of a speaker, and in many similar cases. When this principle is extended to include all sorts of experiences besides those of movement, we have what is generally called Imitation. Moreover, every time that by action the child imitates, he perceives his own imitation, and this again acts as a "copy" or model for another repetition of the act, and so on. This method of keeping himself going gives the young animal or child constant practice, and renders him more and more efficient in the acts necessary to his life.

6. It is evident what great profit accrues from this arrangement whereby a general instinct like imitation takes the place of a number of special instincts, or supplements them. It gives a measure of plasticity to the creature. He can now respond suitably to changes in the environment in which he lives. The special instincts, on the contrary, are for the most part so fixed that the animal must act just as they require him to in this or that circumstance; but as soon as his instinct takes on the form of imitation, the resulting action tends to conform itself to the model actions of the other creatures which set "copies" before him.

These more or less new results due to recent research in the province of Instinct have had direct bearing upon theories of the origin of instinct and of its place in animal life.

Theories of Instinct.—Apart from the older view which saw in animal instinct simply a matter of original created endowment, whereby each animal was made once for all "after his kind," and according to which there is no further reason that the instincts are what they are than that they were made so; apart from this "special creation" view, two different ideas have had currency, both based upon the theory of evolution. Each of these views assumes that the instincts have been developed from more simple animal actions by a gradual process; but they differ as to the elements originally entering into the actions which afterward became instinctive.

1. First, there is what is called the Reflex Theory. This holds that instincts are reflex actions, like the closing of the eye when an object threatens to enter it, only much more complex. They are due to the compounding and adding together of simple reflexes, in greater and greater number, and with increasing efficiency. This theory attempts to account for instinct entirely in terms of nervous action. It goes with that view of evolution which holds that the nervous system has had its growth from generation to generation by the continued reflex adjustments of the organism to its environment, whereby more and more delicate adaptations to the external world were secured. In this way, say the advocates of this theory, we may account for the fact that the animal has no adequate knowledge of what he is doing when he performs an act instinctively; he has no end or aim in his mind; he simply feels his nervous system doing what it is fitted to do by its organic adaptations to the stimulations of air, and earth, and sea, whatever these may be.

But it may be asked: Why do succeeding generations improve each on its parents, so that there is a gradual tendency to perfect the instinct?

The answer to this question brings up another great law of biology—the principle of Variations. This principle states the common fact that in every case of a family of offspring the individual young vary slightly in all directions from their parents. Admitting this, we will find in each group of families some young individuals which are better than their parents; these will have the advantage over others and will be the ones to grow up and have the children of the next generation again, and so on. So by constant Variation and Natural Selection—that is, the "Survival of the Fittest" in competition with the rest—there will be constant improvement in the Instinct.

2. The other theory, the rival one, holds that there are some instincts which show so plainly the marks of Reason that some degree of intelligent adjustment to the environment must be allowed to the animal in the acquiring of these functions. For example, we are told that some of the muscular movements involved in the instincts—such, for example, as the bird's nest-building—are so complex and so finely adjusted to an end, that it is straining belief to suppose that they could have arisen gradually by reflex adaptation alone. There is also a further difficulty with the reflex theory which has seemed insurmountable to many of the ablest psychologists of animal life; the difficulty, namely, that many of the instincts require the action of a great many muscles at the same time, so acting in "correlation" with or support of one another that it is impossible to suppose that the instinct has been acquired gradually. For in the very nature of these cases we can not suppose the instinct to have ever been imperfect, seeing that the partial instinct which would have preceded the perfect performance for some generations would have been not only of no use to the creature, but in many cases positively injurious. For instance, what use to an animal to be able partly to make the movements of swimming, or to the birds to build an inadequate nest? Such instincts would not be usable at all. So we are told by the second theory that the animals must have had intelligence to do these things when they first acquired them. Yet, as is everywhere admitted, after the instinct has been acquired by the species it is then carried out without knowledge and intelligent design, being handed down from generation to generation by heredity.