Speciation of the Wandering Shrew

by James S. Findley

Washington: Whatcom Co.: Blaine, 1 BS; Beaver Creek, 5 WSC; Glacier, 1 BS; Mt. Baker Lodge, 1 WSC; Lake Whatcom, 1 BS; Barron, 2 BS. Okanogan Co.: Sheep Mtn., 3 BS; E. end Bauerman Ridge, 1 BS; Oroville, 1 BS; Hidden Lakes, 1 BS; Loomis, 1 BS; Conconully, 1 BS; Twisp, 1 BS. Ferry Co.: 5 mi. W Curlew, 2 BS. Stevens Co.: Marcus, 1 BS. Pend Oreille Co.: Canyon, 1 WSC; Metaline, 2 BS; Sullivan Lake, 1 BS. San Juan Co.: East Sound, Orcas Island, 3 BS; Friday Harbor, San Juan Island, 1 BS; San Juan Park, 2 WSC; Blakely Island, 1 KU; Richardson, 6 BS. Skagit Co.: Cypress Island, 1 KU; Hamilton, 1 BS; Sauk, 1 BS; Avon, 3 BS; Mt. Vernon, 2 BS; La Conner, 5 BS. Island Co.: San de Fuca, Whidby Island, 3 BS; Greenbank, Whidby Island, 2 BS; 3 mi. N Clinton, Whidby Island, 1 BS. Snohomish Co.: Oso, 2 BS; Hermosa Point, Tulalip Indian Reservation, 7 mi. W and ½ mi. N Marysville, 3 KU. Chelan Co.: Entiat, 2 BS. Lincoln Co.: 6 mi. E Odessa, 4 BS. Spokane Co.: Marshall, 7 BS. Clallam Co.: Neah Bay, 29 BS; 8 mi. W Sekin River, 1 WSC; mouth Sekin River, 1 WSC; Dungeness, 1 BS; Port Townsend, 3 BS; Ozette Indian Reservation, 1 CMNH; Sequim, 4 BS; Tivoli Island, Ozette Lake, 1 CMNH; Garden Island, Ozette Lake, 3 CMNH; Elwah, 1 WSC; Blyn, 1 BS; Soleduck River, 1 BS; 12 mi. S Port Angeles, 1 WSC; Forks, 9 CMNH, 1 BS; Cat Creek, 1 WSC; Lapush, 5 BS. Jefferson Co.: Jefferson Ranger Station, N Fork Hoh River, 5 CMNH; Duckabush, 6 BS. Kitsap Co.: Vashon Island, 2 BS. King Co.: Redmont, 2 BS; Kirkland, 20 BS; Seattle, 1 WSC, 3 KU; Northbend, 2 BS: Lake Washington, near Renton, 2 BS; Kent, 1 BS; Enumclaw, 1 BS. Grays Harbor Co.: Lake Quinault, 9 BS; Aberdeen, 20 BS; Westport, 5 BS, 2 WSC; Oakville, 1 BS. Mason Co.: Lake Cushman, 11 BS; Hoodsport, 1 BS; North Fork Skokomish River, 1 BS; Shelton, 2 BS. Pierce Co.: Puyallup, 6 BS; Steilacoom, 1 BS; 6 mi. S Tacoma, 2 BS; Roy, 3 BS; Bear Prairie, Mt. Rainier, 1 BS; Reflection Lake, Mt. Rainier, 1 WSC. Kittitas Co.: Blewett Pass, 3 BS; Easton, 3 BS; 2 mi. E Cle Elum, 4 FC; Ellensburg, 2 BS. Grant Co.: Moses Lake, 1 BS; 9 mi. S and 1 mi. W Neppel, 1 UM. Whitman Co.: Hangman Creek, Tekoa, 1 WSC; 4 mi. ENE Pullman, 1 KU; 2 mi. N Pullman, 2 WSC; 2 mi. NW Pullman, 1 WSC; 2 mi. W Pullman, 1 WSC; Pullman, 5 WSC; Armstrong, 1 WSC; 5 mi. NE Wawawai, 1 BS; Wawawai, 5 WSC. Thurston Co.: Nisqually Flats, 2 BS; Nisqually, 1 BS; 4 mi. S Olympia, 1 BS; Tenino, 4 BS. Pacific Co.: Tokeland, 4 BS; 1 mi. S Nemah, 2 FC; 1 mi. N Bear River, Willapa Bay, 8 FC; ¼ mi. N Bear River, 3 FC; 3½ mi. E Seaview, 6 FC; Ilwaco, 1 BS. Lewis Co.: 8 mi. W Chehallis, 2 BS; Chehallis, 2 BS; Toledo, 1 BS. Yakima Co.: Selah, 7 KU; Wiley City, 4 BS. Wahkiakum Co.: Cathlamet, 1 BS. Skamania Co.: 45 mi. SE Toledo, 2 BS; Carson, 1 BS; Stevenson, 1 BS; 15 mi. NW White Salmon, 1 BS. Klickitat Co.: Trout Lake, 15 mi. S Mt. Adams, 2 BS; 15 mi. N Goldendale, 1 WSC; Goldendale, 1 BS. Walla Walla Co.: College Place, 1 KU. Columbia Co.: Starbuck, 3 BS. Garfield Co.: 1 mi. E Pomeroy, 1 SGJ. Asotin Co.: 21 mi. SE Dayton, 1 BS; Rogersburg, 1 BS.

Idaho: Bonner Co.: 4 mi. S Sandpoint, 1 UM. Kootenai Co.: Coeur d'Alene, 2 BS. Shoshone Co.: Osburry, 1 BS; Mullan, 2 BS. Latah Co.: Felton's Mills, 1 WSC; Cedar Mtn., 5 WSC. Lewis Co.: Nezperce, 2 BS. Idaho Co.: Seven Devils Mtn., 1 BS. Adams Co.: Summit of Smith Mtn., 7500 ft., 5 KU; New Meadows, 1 BS; Tamarack, 1 BS. Washington Co.: 1 mi. NE Heath, SW slope Cuddy Mtn., 4000 ft., 7 KU. Boise Co.: Bald Mtn. R. S., 10 mi. S Idaho City, 1 BS. Elmore Co.: Cayuse Creek, 10 mi. N Featherville, 1 BS. Canyon Co.: Nampa, 5 BS. Blaine Co.: Sawtooth City, 5 BS; Alturas Lake, 1 BS. Bonneville Co.: 10 mi. SE Irwin, 5 BS. Bannock Co.: Pocatello, 1 BS, 1 KU; 1 mi. W Bancroft, 1 KU; Swan Lake, 1 BS. Owyhee Co.: Grasmere, 1 SGJ. Cassia Co.: 10 mi. S Albion, Mt. Harrison, 1 BS.

Montana: Sanders Co.: Prospect Creek, near Thomson Falls, 4 BS. Lake Co.: Flathead Lake, 5 BS. Ravalli Co.: Bass Creek, NW Stevensville, 2 BS; 2 mi. NE Stevensville, 1 UM; Corvallis, 4 BS; 6 mi. E Hamilton, 1 KU.

Oregon: Clatsop Co.: Seaside, 1 BS. Washington Co.: 5 mi. SE Hillsboro, 1 BS; Beaverton, 1 BS. Multnomah Co.: Portland, 20 BS; Portland, Switzler Lake, 5 BS. Hood River Co.: 2 mi. W Parkdale, 1 BS; north slope Mt. Hood, 2 BS. Umatilla Co.: 10 mi. W Meacham, 2 BS; Meacham, 3 BS. Union Co.: Elgin, 2 BS; Kamela, 2 BS; Hot Lake, 2 BS. Wallowa Co.: 25 mi. N. Enterprise, 4 BS; Wallowa Lake, 23 BS; S Wallowa Lake, 1 BS. Clackamas Co.: Estacada, 1 KU. Marion Co.: Salem, 8 BS; Permilia Lake, 2 BS. Benton Co.: Corvallis, 2 BS; 5 mi. SW Philomath, 5 BS. Linn Co.: Shelburn, 1 BS. Jefferson Co.: 20 mi. W Warm Springs, 2 BS. Grant Co.: Beech Creek, 6 BS; Austin, 1 BS; Strawberry Butte, 1 BS; Strawberry Mts., 12 BS. Baker Co.: Homestead, 1 BS; Cornucopia, 11 BS; Rock Creek, 1 BS; Bourne, 7 BS; McEwen, 1 BS; Huntington, 1 BS; Anthony, 42 AMNH. Lane Co.: north slope Three Sisters, 3 BS; Vida, 1 BS; Mapleton, 1 BS; Eugene, 2 BS; 10 mi. S McKenzie Bridge, 1 BS; Florence, 1 BS. Deschutes Co.: Paulina Lake, 7 BS; Lapine, 8 BS. Crook Co.: 1 SGJ. Douglas Co.: Winchester Bay, 1 SGJ; Scottsburg, 3 BS; Drain, 5 BS; Lookingglass, 1 BS; Diamond Lake, 6 BS. Coos Co.: Empire, 5 BS. Curry Co.: Port Orford, 1 BS; Gold Beach, 4 BS. Klamath Co.: Anna Creek, Mt. Mazama, 1 BS; Crater Lake, 14 BS; Upper Klamath Marsh, 2 BS; Ft. Klamath, 35 BS; Klamath Falls, 6 BS. Lake Co.: 10 mi. SW Silver Lake, 3 BS; west fork Silver Creek, Yamsay Mts., 4 BS; Plush, 1 BS; Warner Creek, Warner Mts., 1 BS; Warner Mts., 3 BS; Gearhart Mts., 17 SGJ; Harney Co.: Diamond, 2 BS; Keiger Gorge, Steens Mts., 3 BS. Malheur Co.: 8 mi. W Jordon Valley, 1 BS.

Wyoming: Lincoln Co.: 13 mi. N and 2 mi. W Afton, 6 KU; 10 mi. N Afton, Salt River, 2 BS; 9 mi. N and 2 mi. W Afton, 6 KU; 7 mi. N and 1 mi. W Afton, 4 KU; Cokeville, 1 BS; 12 mi. N and 2 mi. E Sage, 2 KU; 6 mi. N and 2 mi. E Sage, 1 KU.

California: Del Norte Co.: Smith River, 2 BS; Crescent City, 20 BS. Siskiyou Co.: Beswick, 1 BS; Hornbrook, 3 BS; Brownell, Klamath Lake, 1 BS; Mayten, 2 BS; Squaw Creek, Mt. Shasta, 5 BS; Upper Ash Creek, Mt. Shasta, 1 BS; upper Mud Creek, Mt. Shasta, 8 BS; Wagon Camp, Mt. Shasta, 5 BS; Warmcastle Soda Springs, Squaw Creek Valley, 2 BS; Sisson, 7 BS. Modoc Co.: Davis Creek, Goose Lake, 1 BS. Humboldt Co.: Humboldt Bay, 10 BS. Trinity Co.: Canyon Creek, 2 BS. Shasta Co.: Fort Crook, 11 BS; Dana, 17 BS; Fall Lake, Fall River Valley, 3 BS; Cassel, 2 BS; 12 mi. E Burney, 1 BS; Lassen Peak, 13 BS; Kellys, Warner Creek, 1 KU; Drakes Hot Springs, Warner Creek, 2 BS. Mendocino Co.: Russian Gulch State Park, 2 FC. Plumas Co.: 12 mi. NE Prattville, 2 BS; Spring Garden Ranch, Grizzly Mts., 3 BS; Sierra Valley, 1 BS. Sierra Co.: Lincoln Creek, 1 BS. Sonoma Co.: Petaluma, 3 BS; Point Reyes, 7 BS. Placer Co.: Donner, 3 BS. El Dorado Co.: Tallac, 3 BS. Mono Co.: Mt. Conness, 1 BS; Mono Lake, 1 BS; near Mammoth, 8000 ft., head of Owens River, 2 BS. Inyo Co.: Alvord, 1 BS.

Nevada: Elko Co.: Mountain City, 1 BS; Three Lakes, 1 KU; west side Ruby Lake, 3 mi. N White Pine Co. line, 8 KU; Ruby Mts., 9 BS; W side Ruby Lake, 3 BS. White Pine Co.: W side Ruby Lake, 3 mi. S Elko Co. line, 1 KU. Nye Co.: Cloverdale, Reese River, 3 BS.

Utah: Weber Co.: Beaver Creek, S Fork Ogden River, 2 UU; Huntsville, 10 mi. E Ogden, 1 UU; Hooper Bay Refuge, 4200 ft., 1 UU; Riverdale, 4200 ft., 3 UU; Riverdale, 4250 ft., 1 UU; 3 mi. SE Ogden, 2 UU; Snow Basin, 2 UU; Snow Basin, S part Wheeler Canyon, 1 UU; Uinta, 2 mi. W Weber Canyon entrance, 4 UU; 2 mi. W Uinta, 1 UU. Salt Lake Co.: City Creek Canyon, 6 mi. NE Salt Lake City, 4700 ft., 2 UU; 1 mi. up City Creek Canyon, 4600 ft., 1 UU; ¾ mi. above Forks, City Creek Canyon, 1 UU; The Firs, Millcreek Canyon, 1 UU; Olympus Water Box, 1 UU; Salamander Lake, Lamb's Canyon, 9000 ft., 3 UU (near obscurus); Salt Lake City, 7500 ft., 1 UU; 1 mi. W Draper, 4500 ft., 6 UU; Draper, 4500 ft., 5 UU; 1½ mi. SW Draper, 4500 ft., 1 UU; 3 mi. SW Draper, 4400 ft., 2 UU; 3 mi. S Draper, 4400 ft., 2 UU; 1 mi. S Draper, 4500 ft., 1 UU. Juab Co.: W side Deep Creek Mts., Queen of Sheba Canyon, 8000 ft., 3 UU. Wasatch Co.: Midway Fish Hatchery, 5450 ft., 1 UU.

Marginal records.—British Columbia: Okanagan; Westbridge; Kuskonook; Cranbrook. Montana: Flathead Lake; 6 mi. E Hamilton; Prospect Creek. IDAHO: Cedar Mtn.; New Meadows; Alturas Lake; 10 mi. SE Irwin. Wyoming: 13 mi. N and 2 mi. W Afton; 6 mi. N and 2 mi. E Sage. Idaho: 1 mi. W Bancroft; Swan Lake. Utah: Beaver Creek, South Fork, Ogden River; Midway Fish Hatchery; west side Deep Creek Mts., Queen of Sheba Canyon, 8000 ft. Nevada: Baker Creek (Hall, 1946:120); Reese River (ibid.); 2 mi. S Hinds Hot Springs (ibid.). California: Mono Lake (Jackson, 1928:110); near Mammoth; Alvord; Mount Conness; Donner; Buck Ranch (Jackson, 1928:110); Warner Creek, Drake Hot Springs (ibid.); Canyon Creek; Cuddeback (Jackson, 1928:105); Novato Point (ibid.), thence northward along the coast to Washington: Friday Harbor, San Juan Island. British Columbia: Port Moody.