The cotyledons are very good illustrations of the different uses to which a single organ may be put, and the thorough understanding of it will prepare the pupils' minds for other metamorphoses, and for the theory that all the various parts of a plant are modified forms of a very few members.

4. Nature of the Caulicle.—Probably some of the pupils will have called the caulicle the root. It is, however, of the nature of stem. The root grows only at the end, from a point just behind the tip; the stem elongates throughout its whole length. This can be shown by marking the stem and roots of a young seedling with ink. India ink must be used, as common ink injures the plants. Dip a needle in the ink and prick a row of spots at equal distances on a young root. Corn is very good for this purpose, but Morning-Glory or Bean is better for experiments on the stem. The plants should then be carefully watched and the changes in the relative distance of the spots noted. The experiment is very easily conducted with the seedlings growing on sponge, with their roots in the moist air of the tumbler, as before described.

Dr. Goodale says of this experiment,—"Let a young seedling of corn be grown on damp paper in the manner described in No. 1,[1] and when the longest root is a few centimetres long let it be marked very carefully by means of India ink, or purple ink, put on with a delicate camel's-hair pencil just one centimetre apart. Plants thus marked are to be kept under favorable conditions with respect to moisture and warmth, so that growth will be as rapid as possible. The marks on the older part of the root will not change their relative distance, but the mark at the tip will be carried away from the one next it, showing that the growth has taken place only at this point. Such experiments as the one described are perfectly practicable for all classes of pupils except the very youngest. How far the details of these experiments should be suggested to the pupils, or rather how far they should be left to work out the problem for themselves, is a question to be settled by the teacher in each case. The better plan generally is to bring the problem in a very clear form before the whole class, or before the whole school, and ask whether anybody can think of a way in which it can be solved; for instance, in this case how can it be found out whether roots grow only at their tip or throughout their whole length. If the way is thought out by even a single pupil the rest will be interested in seeing whether the plan will work successfully."

[Footnote 1: Concerning a Few Common Plants, page 25.]

I have been more successful in pricking the roots than in marking them with a brush.

The caulicle can be proved by the manner of its growth to be of the nature of stem, not root. The main root grows from its naked end. Roots can also grow from the sides of the caulicle, as in Indian Corn. In this, it acts precisely as does the stem of a cutting. It can be prettily shown with the seedlings by breaking off a bean at the ground and putting the slip in water. It will throw out roots and the pupil will readily understand that the caulicle does the same thing.

Darwin has made very interesting experiments on the movements of seedlings. If the teacher wishes to repeat some of the experiments he will find the details very fully given in "The Power of Movement of Plants."[1] The pupils can observe in their growing seedlings some of the points mentioned and have already noticed a few in their answers. They have said that the caulicle was the part to grow first, and have spoken of the arched form of the young stem. Their attention should also be drawn to the root-hairs, which are well seen in Corn, Wheat, and Oats. They absorb the liquid food of the plants. A secondary office is to hold the seed firmly, so that the caulicle can enter the ground. This is shown in Red Clover, which may be sown on the surface of the ground. It puts out root-hairs, which attach themselves to the particles of sand and hold the seed. These hairs are treated more fully in the lessons on roots.

[Footnote 1: The Power of Movement in Plants. By Charles Darwin. London. John Murray, 1880.]
[Footnote 1: Reader in Botany. IV. Movements of Seedlings.]

5. Leaves of Seedlings.—Coming now to the question as to the number of leaves at each joint of the stem, the Morning-Glory, Sunflower, and Bean will present no difficulty, but probably all the pupils will be puzzled by the Pea. The stipules, so large and leaf-like, look like two leaves, with a stem between, bearing other opposite leaves, and terminating in a tendril, while in the upper part it could not be told by a beginner which was the continuation of the main stem. For these reasons I left this out in the questions on the Pea, but it should be taken up in the class. How are we to tell what constitutes a single leaf? The answer to this question is that buds come in the axils of single leaves; that is, in the inner angle which the leaf makes with the stem. If no bud can be seen in the Pea, the experiment may be tried of cutting off the top of the seedling plant. Buds will be developed in the axils of the nearest leaves, and it will be shown that each is a compound leaf with two appendages at its base, called stipules, and with a tendril at its apex. Buds can be forced in the same way to grow from the axils of the lower scales, and even from those of the cotyledons, and the lesson may be again impressed that organs are capable of undergoing great modifications. The teacher may use his own judgment as to whether he will tell them that the tendril is a modified leaflet.