Had I had a vote when the Tachytes was christened, I should have suggested a short, harmonious, well-sounding name, meaning nothing else than the thing meant. What better, for example, than the term Sphex? The ear is satisfied and the mind is not corrupted by a prejudice, a source of error to the beginner. I have not nearly as much liking for Ammophila, which represents as a lover of the sands an animal whose establishments call for compact soil. In short, if I had been forced, at all costs, to concoct a barbarous appellation out of Latin or Greek in order to recall the creature's leading characteristic, I should have attempted to say, a passionate lover of the Locust.
Love of the Locust, in the broader sense of the Orthopteron, an exclusive, intolerant love, handed down from mother to daughter with a fidelity which the centuries fail to impair, this, yes, this indeed depicts the Tachytes with greater accuracy than a name smacking of the race-course. The Englishman has his roast-beef; the German his sauerkraut; the Russian his caviare; the Neapolitan his macaroni; the Piedmontese his polenta; the man of Carpentras his tian. The Tachytes has her Locust. Her national dish is also that of the Sphex, with whom I boldly associate her. The methodical classifier, who works in cemeteries and seems to fly the living cities, keeps the two families far removed from each other because of considerations and attaching to the nervures of the wings and the joints of the palpi. At the risk of passing for a heretic, I bring them together at the suggestion of the menu-card.
To my own knowledge, my part of the country possesses five species, one and all addicted to a diet of Orthoptera. Panzer's Tachytes (T. Panzeri, VAN DER LIND), girdled with red at the base of the abdomen, must be pretty rare. I surprise her from time to time working on the hard roadside banks and the trodden edges of the footpaths. There, to a depth of an inch at most, she digs her burrows, each isolated from the rest. Her prey is an adult, medium-sized Acridian (Locust or Grasshopper.—Translator's Note.), such as the White-banded Sphex pursues. The captive of the one would not be despised by the other. Gripped by the antennae, according to the ritual of the Sphex, the victim is trailed along on foot and laid beside the nest, with the head pointing towards the opening. The pit, prepared in advance, is closed for the time being with a tiny flagstone and some bits of gravel, in order to avoid either the invasion of a passer-by or obstruction by landslips during the huntress' absence. A like precaution is taken by the White-banded Sphex. Both observe the same diet and the same customs.
The Tachytes clears the entrance to the home and goes in alone. She returns, puts out her head and, seizing her prey by the antennae, warehouses it by dragging backwards. I have repeated, at her expense, the tricks which I used to play on the Sphex. (For the author's experiments with the Languedocian, the Yellow-winged and the White-edged Sphex, cf. "The Hunting Wasps": chapter 11.—Translator's Note.) While the Tachytes is underground, I move the game away. The insect comes up again and sees nothing at its door; it comes out and goes to fetch its Locust, whom it places in position as before. This done, it goes in again by itself. In its absence I once more pull back the prey. Fresh emergence of the Wasp, who puts things to rights and persists in going down again, still by herself, however often I repeat the experiment. Yet it would be very easy for her to put an end to my teasing: she would only have to descend straightway with her game, instead of leaving it for a moment on her doorstep. But, faithful to the usages of her race, she behaves as her ancestors behaved before her, even though the ancient custom happen to be unprofitable. Like the Yellow-winged Sphex, whom I have teased so often during her cellaring-operations, she is a narrow conservative, learning nothing and forgetting nothing.
Let us leave her to do her work in peace. The Locust disappears underground and the egg is laid upon the breast of the paralysed insect. That is all: one carcase for each cell, no more. The entrance is stopped at last, first with stones, which will prevent the trickling of the embankment into the chamber; next with sweepings of dust, under which every vestige of the subterranean house disappears. It is now done: the Tachytes will come here no more. Other burrows will occupy her, distributed at the whim of her vagabond humour.
A cell provisioned before my eyes on the 22nd of August, in one of the walls in the harmas, contained the finished cocoon a week later. (The harmas was the piece of enclosed waste land in which the author used to study his insects in their natural state. Cf. "The Life of the Fly," by J. Henri Fabre, translated by Alexander Teixeira de Mattos: chapter 1.—Translator's Note.) I have not noted many examples of so rapid a development. This cocoon recalls, in its shape and texture, that of the Bembex-wasps. It is hard and mineralized, this is to say, the warp and woof of silk are hidden by a thick encrustation of sand. This composite structure seems to me characteristic of the family; at all events I find it in the three species whose cocoons I know. If the Tachytes are nearly related to the Spheges in diet, they are far removed from them in the industry of their larvae. The first are workers in mosaic, encrusting a network of silk and sand; the second weave pure silk.
Of smaller size and clad in black with trimmings of silvery down on the edge of the abdominal segments, the Tarsal Tachytes frequents the ledges of soft limestone in fairly populous colonies. (T. tarsina, LEP.) (According to M. J. Perez, to whom I submitted the Wasp of which I am about to speak, this Tachytes might well be a new species, if it is not Lepelletier's T. tarsina or its equivalent, Panzer's T. unicolor. Any one wishing to clear up this point will always recognize the quarrelsome insect by its behaviour. A minute description seems useless to me in the type of investigation which I am pursuing.—Author's Note.) August and September are the season of her labours. Her burrows, very close to one another when an easily-worked vein presents itself, afford an ample harvest of cocoons once the site is discovered. In a certain gravel-pit in the neighbourhood, with vertical walls visited by the sun, I have been able within a short space of time to collect enough to fill the hollow of my hand completely. They differ from the cocoons of the preceding species only in their smaller size. The provisions consist of young Acridians, varying from about a quarter to half an inch in length. The adult insect does not appear in the assorted bags of game, being no doubt too tough for the feeble grub. All the carcases consist of Locust-larvae, whose budding wings leave the back uncovered and put one in mind of the short skirts of a skimpy jacket. Small so that it may be tender, the game is numerous so that it may suffice all needs. I count from two to four carcases to a cell. When the time comes we will discover the reason for these differences in the rations served.
The Mantis-killing Tachytes wears a red scarf, like her kinswoman, Panzer's Tachytes. (The Mantis-hunting Tachytes was submitted to examination by M. J. Perez, who failed to recognize her. This species may well be new to our fauna. I confine myself to calling her the Mantis-killing Tachytes and leave to the specialists the task of adorning her with a Latin name, if it be really the fact that the Wasp is not yet catalogued. I will be brief in my delineation. To my thinking the best description is this: mantis-hunter. With this information it is impossible to mistake the insect, in my district of course. I may add that it is black, with the first two abdominal segments, the legs and the tarsi a rusty red. Clad in the same livery and much smaller than the female, the male is remarkable for his eyes, which are of a beautiful lemon-yellow when he is alive. The length is nearly half an inch for the female and a little more than half this for the male.—Author's Note.) I do not think that she is very widely distributed. I made her acquaintance in the Serignan woods, where she inhabits, or rather used to inhabit—for I fear that I have depopulated and even destroyed the community by my repeated excavations—where she used to inhabit one of those little mounds of sand which the wind heaps up against the rosemary clumps. Outside this small community, I never saw her again. Her history, rich in incident, will be given with all the detail which it deserves. I will confine myself for the moment to mentioning her rations, which consist of Mantis-larvae, those of the Praying Mantis predominating. (Cf. "The Life of the Grasshopper": chapters 6 to 9.—Translator's Note.) My lists record from three to sixteen heads for each cell. Once again we note a great inequality of rations, the reason for which we must try to discover.
What shall I say of the Black Tachytes (T. nigra, VAN DER LIND) that I have not already said in telling the story of the Yellow-winged Sphex? ("The Hunting Wasps": chapters 4 to 6.—Translator's Note.) I have there described her contests with the Sphex, whose burrow she seems to me to have usurped; I show her dragging along the ruts in the roads a paralysed Cricket, seized by the hauling-ropes, his antennae; I speak of her hesitations, which lead me to suspect her for a homeless vagabond, and finally on her surrender of her game, with which she seems at once satisfied and embarrassed. Save for the dispute with the Sphex, an unique event in my records as observer, I have seen all the rest many a time, but never anything more. The Black Tachytes, though the most frequent of all in my neighbourhood, remains a riddle to me. I know nothing of her dwelling, her larvae, her cocoons, her family-arrangements. All that I can affirm, judging by the invariable nature of the prey which one sees her dragging along, is that she must feed her larvae on the same non-adult Cricket that the Yellow-winged Sphex chooses for hers.
Is she a poacher, a pillager of other's property, or a genuine huntress? My suspicions are persistent, though I know how chary a man should be of suspicions. At one time I had my doubts about Panzer's Tachytes, whom I grudged a prey to which the White-banded Sphex might have laid claim. To-day I have no such doubts: she is an honest worker and her game is really the result of her hunting. While waiting for the truth to be revealed and my suspicions set aside, I will complete the little that I know of her by noting that the Black Tachytes passes the winter in the adult form and away from her cell. She hibernates, like the Hairy Ammophila. In warm, sheltered places, with low, perpendicular, bare banks, dear to the Wasps, I am certain of finding her at any time during the winter, however briefly I investigate the earthen surface, riddled with galleries. I find the Tachytes cowering singly in the hot oven formed by the end of a tunnel. If the temperature be mild and the sky clear, she emerges from her retreat in January and February and comes to the surface of the bank to see whether spring is making progress. When the shadows fall and the heat decreases, she reenters her winter-quarters.