But there is a lizard more interesting still, and that is the crocodile. He stands alone among reptiles. His heart has two ventricles, and you would think that he ought to be included in the class of warm-blooded animals. But, no. The separation of the two kinds of blood takes place in the heart, it is true, and it is really true arterial blood which the aorta carries away from the left ventricle. But the right ventricle has two doors of exit. One communicates with the lungs, the other with the aorta; and the latter has hardly performed its distribution in the upper part of the body when it meets, as it descends, with a treacherous tube bringing to it a current of venous blood. In this way only half the blood that comes from the veins passes on to be regenerated by contact with the air, and all the hinder part of the body receives nothing but the mixed blood common to reptiles, while the head and fore members enjoy the privilege of the superior orders. After this go and lay down your laws of classification! Nature, while maintaining amongst all animals the same principle of life—the regeneration of the blood by oxygen—has in their construction followed many systems leading to the same result by different combinations, and which seem to permit the establishment of essential distinctions among them. Here is an animal who, if I may so express myself, is climbing up from one system to the other, and you would have to cut him in two before you could classify him properly, since his fore-quarters have risen to the warm-blooded animals, while his hind ones are left behind among the cold-blooded reptiles!
But there is something which even outdoes this.
On dry land the crocodile is timid, faltering, a bad walker, incapable of regular combat, and a man can manage him with a stick. One feels that he is betrayed by the hinder half of his body, through which circulates the only half-oxygenized blood. But when once he has plunged into the water his whole behavior suddenly alters; he is a ferocious being, high-mettled, indomitable, a savage enemy, redoubling his exertions, as if the entire mass of his blood had suddenly become arterial. Geoffroy St. Hilaire, who followed Bonaparte as a scientific explorer when he set out for the conquest of Egypt, the country of crocodiles, was deeply struck by studying on the spot this double life, which seems in a way to maintain two beings in the same body. He afterwards gave an extremely curious explanation of it in his work on the crocodiles of Egypt. Here it is; but I forewarn you that you will not understand it:
"The crocodile, when it is under water, receives by two canals into the cavity of the abdomen, a considerable quantity of water, which the animal can renew at will."
You are not much the wiser, are you? But wait a moment. We are soon coming to the fishes, and you will then see what an unlimited scope nature has allowed herself here. Not satisfied with two systems in one animal, she appears to have got hold of three.
If we continue the examination of this privileged reptile, we shall find many other infractions of the usual rules of his class. His tongue, certainly, is fastened to his mouth like that of the tortoise, so much so that the ancient Egyptians told the Greeks he had not got one; but his set of teeth clearly approach those of the lower mammals. You have probably heard a great deal of the strength of the crocodile's formidable teeth. Travellers have given them this reputation; but we have nothing to do with that now. They stand in battle array, in a single line, along the whole length of the jaws, into which they are sunk with genuine fangs, whilst the prongs of our little lizard are merely fastened to the surface of the bones which support them. Indeed, in one way, the crocodile is even better provided than the mammals. He possesses under each tooth one or two germs, the life of which lasts as long as that of the animal, and which are always there ready toreplace the previous one should it chance to fall out. There are many ladies, and (not to be rude) gentlemen as well, who would, I am sure, give a great deal to have as many teeth at their service. Indeed, they may possibly think Dame Nature very unjust to have selected this great villanous beast rather than us as the object of a gift which they would have been so well able to appreciate. But we must not blame nature too quickly: she had her reasons. We, during our infancy, have teeth in reserve. Now, a reptile may be considered as an imperfect rough draft of a mammal; and the crocodile gives one thoroughly the idea of a mammal half-finished and fixed for life in a state of childhood. I am sorry that I cannot enter into full details, that you might see how far the idea is a just one. Moreover, in his character of a perpetual child, he is always growing bigger all his life long, and never seems able to die but by accident, hardly ever, I may really say, from old age. By specimens kept in captivity, it has been ascertained that their growth is very slow. Well, imagine their being only from seven to eight inches long when they come out of the shell, and that full-grown crocodiles have been found thirty feet in length, and calculate accordingly. You will not account for it under a century; and I should like to know what would become of this venerable child of more than a hundred years old if kind Mother Nature had not left him our system of milk-teeth to the end?
A curious peculiarity of these persistent teeth is, that they are hollow inside, so much so, that the bowls of tobacco-pipes are said to be made from them in Europe. I mention the fact, although of no great interest to you, for the benefit of any pipe-merchants who have not yet thought of sending for such things to Cairo.
But let us return to the efforts perceptible in the organization of the crocodile to raise itself to a higher level. The soft palate, as we called it (Letter VII.), is wanting in other reptiles; but here there is one which completely closes the entrance of the windpipe (the larynx). I announced, too, the disappearance of the diaphragm; and we bewailed together the loss of that servant of the good old times, whose touching history you must, I am sure, remember. But I reckoned without this wretched crocodile, who seems determined to give the lie to all we have been saying. He has a diaphragm, and one which acts well enough in the main, although it is pierced right through the middle, as if it were rather ashamed of being there, and wished to make up for dividing the body into two compartments, against all proper reptile regulations, by opening a door of communication between them. What shall I tell you besides? The lungs, not to be behind the rest of this aristocratic reptile's organization, are hollowed into cells much more complicated than those of his fellows. You find here no end of nooks and corners, which multiply opportunities of contact between the air and the blood, and so give the crocodile almost the respiration of the mammals, as he has already got pretty nearly their system of circulation.
With serpents, again, we fall very low. When we were speaking of the tortoises I told you that, in proportion as we come down in the scale, the digestive tube has a tendency to get rid of its accessories, and to assume the appearance of a perfectly straight tube. If any one were to cut open a serpent before you, you would see this final condition almost reached already. In the first place, the soft palate is entirely suppressed, and the mouth extends straight into the oesophagus, whose tube seems to run through the whole length of the body without interruption, with just four or five doublings towards the base, in that part which represents the intestines. An imperceptible swelling indicates the place where the real stomach lies within; but in another sense one may call the oesophagus, and I might almost add the mouth itself, its stomach. You shall see how.
The jaws of serpents are even in a more unfinished state than those of other reptiles. Nature has not taken the time to weld the different parts of them together; but these begin by not being very firmly joined, remember, in young mammals. The bones of the head, which support the jaws, are themselves movable, and can be detached from the skull if necessary, so as to allow the throat to open extraordinarily wide; thus it is not uncommon to see a serpent swallow animals much larger than itself. You will be horrified when I tell you that the anaconda, one of the giants of the family, swallows large quadrupeds at a single mouthful. What are our mouthfuls in comparison with his? however, it must be confessed, that his often take several days to go down. When the animal has rolled up his prey in his terrible folds, he pounds and kneads it till it is reduced to a kind of long roll, which he moistens with a copious slaver to make it slip down more easily. Then, attacking it at one end, he fastens this very expansive jaw upon it, and the gigantic mouthful slowly begins its journey; what was left outside the mouth, advancing little by little, in proportion as the digestion reduces what has entered to pulp, and sends it farther down. This is on great occasions; but in the case of more modest prey—a rabbit, for instance—the mouthful goes in whole at one gulp and remains stationary, partly in the oesophagus, partly in the stomach, while the powerful juices distilled by the walls of the latter are dissolving it.