Dry-Farming : A System of Agriculture for Countries under a Low Rainfall

by John Andreas Widtsoe

It is clear that under these circumstances not only water enters the root-hairs, but many of the substances found in solution in the soil-water enter the plant also. Among these are the mineral substances which are indispensable for the proper life and growth of plants. These plant nutrients are so indispensable that if any one of them is absent, it is absolutely impossible for the plant to continue its life functions. The indispensable plant-foods gathered from the soil by the root-hairs, in addition to water, are: potassium, calcium, magnesium, iron, nitrogen, and phosphorus,—all in their proper combinations. How the plant uses these substances is yet poorly understood, but we are fairly certain that each one has some particular function in the life of the plant. For instance, nitrogen and phosphorus are probably necessary in the formation of the protein or the flesh-forming portions of the plant, while potash is especially valuable in the formation of starch.

There is a constant movement of the indispensable plant nutrients after they have entered the root-hairs, through the stems and into the leaves. This constant movement of the plant-foods depends upon the fact that the plant consumes in its growth considerable quantities of these substances, and as the plant juices are diminished in their content of particular plant-foods, more enters from the soil solution. The necessary plant-foods do not alone enter the plant but whatever may be in solution in the soil-water enters the plant in variable quantities. Nevertheless, since the plant uses only a few definite substances and leaves the unnecessary ones in solution, there is soon a cessation of the inward movement of the unimportant constituents of the soil solution. This process is often spoken of as selective absorption; that is, the plant, because of its vital activity, appears to have the power of selecting from the soil certain substances and rejecting others.

Movement of water through plant

The soil-water, holding in solution a great variety of plant nutrients, passes from the root-hairs into the adjoining cells and gradually moves from cell to cell throughout the whole plant. In many plants this stream of water does not simply pass from cell to cell, but moves through tubes that apparently have been formed for the specific purpose of aiding the movement of water through the plant. The rapidity of this current is often considerable. Ordinarily, it varies from one foot to six feet per hour, though observations are on record showing that the movement often reaches the rate of eighteen feet per hour. It is evident, then, that in an actively growing plant it does not take long for the water which is in the soil to find its way to the uppermost parts of the plant.

The work of leaves

Whether water passes upward from cell to cell or through especially provided tubes, it reaches at last the leaves, where evaporation takes place. It is necessary to consider in greater detail what takes place in leaves in order that we may more clearly understand the loss due to transpiration. One half or more of every plant is made up of the element carbon. The remainder of the plant consists of the mineral substances taken from the soil (not more than two to 10 per cent of the dry plant) and water which has been combined with the carbon and these mineral substances to form the characteristic products of plant life. The carbon which forms over half of the plant substance is gathered from the air by the leaves and it is evident that the leaves are very active agents of plant growth. The atmosphere consists chiefly of the gases oxygen and nitrogen in the proportion of one to four, but associated with them are small quantities of various other substances. Chief among the secondary constituents of the atmosphere is the gas carbon dioxid, which is formed when carbon burns, that is, when carbon unites with the oxygen of the air. Whenever coal or wood or any carbonaceous substance burns, carbon dioxid is formed. Leaves have the power of absorbing the gas carbon dioxid from the air and separating the carbon from the oxygen. The oxygen is returned to the atmosphere while the carbon is retained to be used as the fundamental substance in the construction by the plant of oils, fats, starches, sugars, protein, and all the other products of plant growth.

This important process known as carbon assimilation is made possible by the aid of countless small openings which exist chicfly on the surfaces of leaves and known as "stomata." The stomata are delicately balanced valves, exceedingly sensitive to external influences. They are more numerous on the lower side than on the upper side of plants. In fact, there is often five times more on the under side than on the upper side of a leaf. It has been estimated that 150,000 stomata or more are often found per square inch on the under side of the leaves of ordinary cultivated plants. The stomata or breathing-pores are so constructed that they may open and close very readily. In wilted leaves they are practically closed; often they also close immediately after a rain; but in strong sunlight they are usually wide open. It is through the stomata that the gases of the air enter the plant through which the discarded oxygen returns to the atmosphere.

It is also through the stomata that the water which is drawn from the soil by the roots through the stems is evaporated into the air. There is some evaporation of water from the stems and branches of plants, but it is seldom more than a thirtieth or a fortieth of the total transpiration. The evaporation of water from the leaves through the breathing-pores is the so-called transpiration, which is the greatest cause of the loss of soil-water under dry-farm conditions. It is to the prevention of this transpiration that much investigation must be given by future students of dry-farming.

Transpiration

As water evaporates through the breathing-pores from the leaves it necessarily follows that a demand is made upon the lower portions of the plant for more water. The effect of the loss of water is felt throughout the whole plant and is, undoubtedly, one of the chief causes of the absorption of water from the soil. As evaporation is diminished the amount of water that enters the plants is also diminished. Yet transpiration appears to be a process wholly necessary for plant life. The question is, simply, to what extent it may be diminished without injuring plant growth. Many students believe that the carbon assimilation of the plant, which is fundamentally important in plant growth, cannot be continued unless there is a steady stream of water passing through the plant and then evaporating from the leaves.