Fig. 8: Syrrhophus rubrimaculatus (upper right, KU 58911, ×1.6; lower right, KU 58910, ×4) and S. verrucipes (upper left, UIMNH 15995, ×1.6; lower left, UIMNH 15989, ×3.7).
Diagnosis.—Small frogs, males 18.2-23.5 mm. snout-vent, females 19.0-22.5 mm. snout-vent length (small sample); vocal slits in males; digital tips scarcely expanded ([Fig. 1]); first finger shorter than second; outer palmar tubercle reduced in size; inner metatarsal tubercle elongate, twice the size of small, conical outer metatarsal tubercle; diameter of tympanum 35.5-46.5 per cent that of eye in both sexes; dorsum brown with small pale spots (red in life); venter gray.
Remarks.—Previous authors who treated Syrrhophus placed this species in the western complex, because it occurs on the Pacific versant and has a reduced outer palmar tubercle. Duellman (1958) placed rubrimaculatus apart from the other western species, because of its relatively unexpanded digital tips and coloration. The digital tips are like those in leprus, which rubrimaculatus resembles. Except for the reduction of the outer palmar tubercle, rubrimaculatus could be a member of the leprus group.
Syrrhophus rubrimaculatus is probably best treated as a Pacific derivative of the leprus group, even though the palmar tubercles do not agree. The removal of rubrimaculatus from the western complex results in a more homogeneous remainder and does not greatly increase the heterogeneity of the eastern complex.
Etymology.—Latin, meaning spotted with red; in reference to the colors in life.
Distribution.—Low to moderate elevations on the Pacific versant of southeastern Chiapas, México ([Fig. 7]); probably extending into adjacent Guatemala.
Specimens examined.—(48) MÉXICO, Chiapas: Escuintla, UMMZ 88283; 6 km. NE Escuintla, UMMZ 87876-80; La Esperanza, UIMNH 13285, UMMZ 88496-97, USNM 114070 (holotype), 114054-69, 114072; Monte Cristo, UMMZ 88353; 1.3 km. N Puerto Madero, KU 58910-11; Finca San Jerónimo, 600-650 m., UIMNH 55299-312, 55313-16 (cleared and stained).
Malachylodes guttilatus Cope, 1879:264 [Holotype.—USNM 9888, from Guanajuato, Guanajuato, México; collected in 1877 by Alfredo Duges].