The physiological action is in this case clear. The vibration of the nerve caused by the tickling travels from the foot to the appropriate centre in the spinal marrow, and here gives rise to, or is switched off as, a motor impulse travelling back to the muscles of the leg, causing them to contract. In the injured patient the nervous impulse cannot reach the brain, the seat of consciousness, and hence this is not awakened. Normally consciousness does result in a majority of such cases, but only after the beginning or completion of the appropriate action. Yet the movements of our internal organs, intestine and heart, go on continually, and in health we remain entirely unconscious of their action.
But reflex actions may be anything but simple. We walk and talk, and write or play the piano without ever thinking of a single muscle or organ. Yet we had once to learn with much effort to take each step or frame each letter. Thus actions, originally conscious and intended, easily become reflex; often repeated the brain leaves their control to the lower centres. We often say, "I did not intend to do that; I could not help it." We forget that this excuse is our worst condemnation. It is a confession that we have allowed or encouraged a habit to wear a groove from which the wheels of our life cannot escape. The essential characteristic of reflex action is therefore that from beginning to completion it goes on independently of consciousness.
2. Instinct. This is a much-abused word. It is frequently applied to all the mental actions of animals without much thought or care as to its meaning. Let us gain a definition from the study of a typical case lest we use the word as a cloak for ignorance or negligent thoughtlessness. Watch a spider building its wonderful geometrical web. The web is a work of art, and every motion of the spider beautifully adapted to its purpose. But the spider is not therefore necessarily an artist. Let us see of how much the spider is probably conscious, remembering that our best judgment is but an inference. We have good reason to believe that she is conscious of the stimulus to action, hunger. She may be, probably is, conscious of the end to be attained—to catch a fly for her dinner. She seems conscious of what she is doing. In all these respects this differs from reflex action. But she is probably unconscious of the exact fitness of the means to the end. We do not believe that she has adopted the geometrical pattern, because she has discovered or calculated that this will make the closest and largest net for the smallest outlay of labor and material. Furthermore the young spider builds practically as good a web as the old one. She has inherited the power, not developed or gained it by experience or observation. And all the members of the species have inherited it in much the same degree of perfection.
Concerning the origin of instincts there are several theories. Some instincts would seem to be the result of non-intelligent, perhaps unconscious, habits becoming fixed by heredity and improved by natural selection; others would appear to be modifications of actions originally due to intelligence. Instinct is therefore characterized by consciousness of the stimulus to act, of the means and end, without the knowledge of the exact adaptation of means to end. It is hereditary and characterizes species or large groups.
3. Intelligent Action. You come in cold and sit down before an open fire. You push the brands together to make the fire burn. Applying once more the criterion of consciousness to this action we notice that you are conscious of the stimulus to act, of the steps of the action, and of the end to be attained, exactly as in instinctive action. But finally, and this is the essential characteristic of intelligent action, you are aware to a certain extent of the fitness of the means to the attainment of the end. This piece of knowledge you had to acquire for yourself. Erasmus Darwin defined a fool as a man who had never tried an experiment. Experience and observation, not heredity, are the sources of intelligence. Intelligence is power to think, and a man may be very learned—for do we not have learned pigs?—and yet have very little real intelligence. Hence this is possessed by different individuals in very varying degrees.
We may now briefly compare these three kinds of nervous action.
Reflex action is involuntary and unconscious. The actor may, and usually does, become conscious of the action after it has been commenced or completed, but this is not at all necessary or universal.
Instinctive action is to a certain extent voluntary and conscious. The actor is conscious of the stimulus, the means and mode, and the end or purpose of the action. Of the exact fitness or adaptation of the means to the end the actor is unconscious.
Intelligent action is conscious and voluntary. The actor is conscious of the stimulus to act, of the means and mode, and to a certain extent of the adaptation of the means to the end. This last item of knowledge, lacking in instinctive action, is acquired by experience or observation.
Reflex action may be regarded as a comparatively mechanical, though often very complex, process; the reflex ganglia appear to be hardly more than switch-boards. There is stimulus of the sense-organs, and thus what Mr. Romanes has called "unfelt sensation," unfelt as far as the completion of the action is concerned. But in instinct the sensation no longer remains unfelt; perception is necessary, consciousness plays a part. And this consciousness is a vastly more subtle element, differing as much apparently from the vibration of brain, or nervous, molecules as the Geni from the rubbing of Aladdin's lamp, to borrow an illustration.