26. I have looked wholly in vain through all my books on birds, to find some account of the muscles it uses in hopping, and of the part of the toes with which the spring is given. I must leave you to find out that for yourselves; it is a little bit of anatomy which I think it highly desirable for you to know, but which it is not my business to teach you. Only observe, this is the point to be made out. You leap yourselves, with the toe and ball of the foot; but, in that power of leaping, you lose the faculty of grasp; on the contrary, with your hands, you grasp as a bird with its feet. But you cannot hop on your hands. A cat, a leopard, and a monkey, leap or grasp with equal ease; but the action of their paws in leaping is, I imagine, from the fleshy ball of the foot; while in the bird, characteristically γαμφωνυξ [Greek: gampsônux], this fleshy ball is reduced to a boss or series of bosses, and the nails are elongated into sickles or horns; nor does the springing power seem to depend on the development of the bosses. They are far more developed in an eagle than a robin; but you know how unpardonably and preposterously awkward an eagle is when he hops. When they are most of all developed, the bird walks, runs, and digs well, but leaps badly.

27. I have no time to speak of the various forms of the ankle itself, or of the scales of armor, more apparent than real, by which the foot and ankle are protected. The use of this lecture is not either to describe or to exhibit these varieties to you, but so to awaken your attention to the real points of character, that, when you have a bird's foot to draw, you may do so with intelligence and pleasure, knowing whether you want to express force, grasp, or firm ground pressure, or dexterity and tact in motion. And as the actions of the foot and the hand in man are made by every great painter perfectly expressive of the character of mind, so the expressions of rapacity, cruelty, or force of seizure, in the harpy, the gryphon, and the hooked and clawed evil spirits of early religious art, can only be felt by extreme attention to the original form.

28. And now I return to our main question, for the robin's breast to answer, "What is a feather?" You know something about it already; that it is composed of a quill, with its lateral filaments terminating generally, more or less, in a point; that these extremities of the quills, lying over each other like the tiles of a house, allow the wind and rain to pass over them with the least possible resistance, and form a protection alike from the heat and the cold; which, in structure much resembling the scale-armor assumed by man for very different objects, is, in fact, intermediate, exactly, between the fur of beasts and the scales of fishes; having the minute division of the one, and the armor-like symmetry and succession of the other.

29. Not merely symmetry, observe, but extreme flatness. Feathers are smoothed down, as a field of corn by wind with rain; only the swathes laid in beautiful order. They are fur, so structurally placed as to imply, and submit to, the perpetually swift forward motion. In fact, I have no doubt the Darwinian theory on the subject is that the feathers of birds once stuck up all erect, like the bristles of a brush, and have only been blown flat by continual flying.

Nay, we might even sufficiently represent the general manner of conclusion in the Darwinian system by the statement that if you fasten a hair-brush to a mill-wheel, with the handle forward, so as to develop itself into a neck by moving always in the same direction, and within continual hearing of a steam-whistle, after a certain number of revolutions the hair-brush will fall in love with the whistle; they will marry, lay an egg, and the produce will be a nightingale.

30. Whether, however, a hog's bristle can turn into a feather or not, it is vital that you should know the present difference between them.

The scientific people will tell you that a feather is composed of three parts—the down, the laminæ, and the shaft.

But the common-sense method of stating the matter is that a feather is composed of two parts, a shaft with lateral filaments. For the greater part of the shaft's length, these filaments are strong and nearly straight, forming, by their attachment, a finely warped sail, like that of a wind-mill. But towards the root of the feather they suddenly become weak, and confusedly flexible, and form the close down which immediately protects the bird's body.

To show you the typical arrangement of these parts, I choose, as I have said, the robin; because, both in his power of flying, and in his color, he is a moderate and balanced bird;—not turned into nothing but wings, like a swallow, or nothing but neck and tail, like a peacock. And first for his flying power. There is one of the long feathers of robin's wing, and here (Fig. 1) the analysis of its form.

31. First, in pure outline (A), seen from above, it is very nearly a long oval, but with this peculiarity, that it has, as it were, projecting shoulders at a 1 and a 2. I merely desire you to observe this, in passing, because one usually thinks of the contour as sweeping unbroken from the root to the point. I have not time to-day to enter on any discussion of the reason for it, which will appear when we examine the placing of the wing feathers for their stroke.