In November 1832, Darwin returned to Monte Video, from a long journey in the interior of the South American Continent, bringing with him many zoological specimens and a great quantity of fossil bones, teeth and scales, dug out by him with infinite toil from the red mud of the Pampas—these fossils evidently belonging to the geological period that immediately preceded that of the existing creation. The living animals represented in his collection were all obviously very distinct from those of Europe—consisting of curious sloths, anteaters, and armadilloes—the so-called 'Edentata' of naturalists. And when young Darwin came to examine and compare his fossil bones, teeth and scales he found that they too must have belonged to animals (megatherium, mylodon, glyptodon, etc.) quite distinct from but of strikingly similar structure to those now living in South America. What could be the meaning of this wonderful analogy? If Cuvier and his fellow Catastrophists were correct in their view that, at each 'revolution' taking place on the earth's surface, the whole batch of plants and animals was swept out of existence, and the world was restocked with a 'new creation,' why should the brand-new forms, at any particular locality, have such a 'ghost-like' resemblance to those that had gone before? It is interesting to note that, just at the same time, a similar discovery was made with respect to Australia. In caves in that country, a number of bones were found which, though evidently belonging to 'extinct' animals, yet must have belonged to forms resembling the kangaroos and other 'pouched animals' (marsupials) now so distinctive of that continent. But of this fact Darwin was not aware until after his return to England in 1836.

Among the objects sent from home, which awaited Darwin on his return to Monte Video, was the second volume of Lyell's Principles, then newly published; this book, while rejecting Lamarckism, was crowded with facts and observations concerning variation, hybridism, the struggle for existence, and many other questions bearing on the great problem of the origin of species. I think there can be no doubt that from this time Darwin came to regard the question of species with an interest he had never felt before.

It is of course not suggested that, at this early date, Darwin had formed any definite ideas as to the mode in which new species might possibly arise from pre-existing ones or even that he had been converted to a belief in evolution. Indeed in 1877 he wrote 'When I was on board the Beagle I believed in the permanence of species' yet he adds 'but as far as I can remember vague doubts occasionally flitted across my mind.' Such 'vague doubts' could scarcely have failed to have arisen when, as happened during all his journeys from north to south of the South American Continent, he found the same curious correspondence between existing and late fossil forms of life again and again illustrated.

But towards the end of the voyage, an even stronger element of doubt as to the immutability of species was awakened in his mind. When he came to study the forms of life existing in the Galapagos Islands, off the west coast of South America, he was startled by the discovery of the following facts. Each small island had its own 'fauna' or assemblage of animals—this being very strikingly shown in the case of the reptiles and birds. And yet, though the species were different, there was obviously a very wonderful 'family likeness' to one another between the forms in the several islands and between them all and the animals living in the adjoining portion of the continent. Surely this could not be accidental, but must indicate relationships due to descent from common ancestors!

Charles Darwin returned to England in 1836, and at once made the acquaintance of Lyell. He says in one place, 'I saw a great deal of Lyell' and in another that 'I saw more of Lyell than of any other man, both before and after my marriage.' In one of his letters he writes, 'You cannot conceive anything more thoroughly good natured than the heart-and-soul manner in which he put himself in my place and thought what would be best to do[105].' For two years Darwin was comparatively free from the distressing malady which clouded so much of his after life. And, during that time, he engaged very heartily with Lyell in those combats at the Geological Society (of which he had become one of the Secretaries) in which their joint views concerning the truth of continuity or evolution in the inorganic world were defended against the attacks of the militant catastrophists. Darwin, however, did not act on the defensive alone, but brought forward a number of papers strongly supporting his new friend's views.

There can be little doubt that, while thus engaged, and in constant friendly intercourse with Lyell, Darwin must have felt—like other earnest thinkers on geology at that day—that the principles they were advocating of 'continuity' in the inorganic world must be equally applicable to the organic world—and thus that the question of evolution would acquire a new interest for him.

But it was undoubtedly the revision of the notes made on board the Beagle, and the study of the specimens which had been sent home by him from time to time, that produced the great determining influence on Darwin's career. All through the voyage he had endeavoured, with as much literary skill as he could command, to record with accuracy the observations he made, and the conclusions to which, on careful reflection, they seemed to point. And on his return to England, these patiently written journals were revised and prepared for publication forming that charming work A Naturalist's Voyage. Journal of Researches into the Natural History and Geology of the Countries visited during the Voyage of H.M.S. 'Beagle' round the world.

As Darwin, with the specimens before him, revised his notes, and reconsidered the impressions made on his mind, the 'vague doubts' he had entertained, from time to time, concerning the immutability of species, would come back to him with new force and cumulative effect. 'I then saw,' he says, 'how many facts indicated the common descent of species,' and further, 'It occurred to me in 1837, that something might perhaps be made out on this question by patiently accumulating and reflecting on all sorts of facts which could possibly have any bearing on it.' In July of that year, he opened his first note-book on the subject[106]—the note-books being soon replaced by a series of portfolios, in which extracts from the various works he read, facts obtained by correspondence, the records of experiments and observation, and ideas suggested by constant meditation were slowly accumulated for twenty years. Mr Francis Darwin has published a series of extracts from the note-book of 1837, which amply prove that by this time Charles Darwin had become 'a convinced evolutionist[107].'

Fifteen months after this 'systematic enquiry' began, Darwin happened to read the celebrated work of Malthus On Population, for amusement, and this served as a spark falling on a long prepared train of thought. The idea that as animals and plants multiply in geometrical progression, while the supplies of food and space to be occupied remain nearly constant, and that this must lead to a 'struggle for existence' of the most desperate kind, was by no means new to Darwin, for the elder De Candolle, Lyell and others had enlarged upon it; yet the facts with regard to the human race, so strikingly presented by Malthus, brought the whole question with such vividness before him that the idea of 'Natural Selection' flashed upon Darwin's mind. This hypothesis cannot be better or more succinctly stated than in Huxley's words.

'All species have been produced by the development of varieties from common stocks: by the conversion of these, first into permanent races and then into new species, by the process of natural selection, which process is essentially identical with that artificial selection by which man has originated the races of domestic animals—the struggle for existence taking the place of man, and exerting, in the case of natural selection, that selective action which he performs in artificial selection[108].'