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Key to Species and Subspecies

1. Larger frogs (males to 49 mm. snout-vent length); thighs strongly barred; supratympanic fold black; dorsum blotched or spotted2

Smaller frogs (males to 40 mm. snout-vent length); thighs weakly barred or plain; supratympanic fold pale brown; dorsum usually having linear pattern3

2. Dorsum tuberculate; snout subacuminate; vocal sac flecked with brown; tarsal fold rudimentary; web absent between fingers; black spots absent in scapular regionH. boulengeri

Dorsum smooth; snout pointed; vocal sac dark gray; tarsal fold absent; trace of web between fingers; two or more elongate dark spots in scapular regionH. foliamorta

3. Snout-vent length more than 30 mm.; tympanum 2/3 to ¾ diameter of eye; prevomerine elevations about size of choanae4

Snout-vent length less than 30 mm.; tympanum less than ½ diameter of eye; prevomerine elevations smaller than choanae5

4. Thighs mottled posteriorly; discs on fingers about ½ size of tympanum; faint dark line from nostril to eyeH. rubra

Thighs faintly barred or plain posteriorly; discs on fingers about size of tympanum; distinct dark line from nostril to eye H. elaeochroa