Questions of fact are simple to deal with. It is indubitable that some forms of life remain stationary and unevolving for secular periods; it is equally indubitable that degeneration is widespread in evolution. These are facts. But we are not therefore called upon to deny the possibility of progress. To do so would be to fall into the error of reasoning which we have already condemned. It remains for us to take these facts into account when examining the totality of facts concerning organic life, and to see whether, in spite of them, we cannot discover a series of other facts, a movement in phenomena, which may still legitimately be called progress. To deny progress because of degeneration is really no more legitimate than to assert that, because each wave runs back after it has broken, therefore the tide can never rise.
Similarly with the first two objections. If the degree of adaptation has not increased during evolution, then it is clear that progress does not consist in increase in adaptation. But it does not follow that progress does not exist; it may quite well consist in an increase of other qualities. So with complexity. Complexity has increased, but increase in complexity is not progress, say the objectors. Granted: but may there not be something else which has increased besides mere complexity?
No; the remedy for all our difficulties, and indeed the only way in which we can arrive at the possibility of saying whether biological progress exists or no, is to adopt the positive method.
Let us then begin our survey of biological evolution in the endeavour to find whether or no progress is visible there. To start with, we must be clear what are the sources of our knowledge on the subject.
Direct observation of progressive evolution has, of course, not yet been possible in the period—biologically negligible—in which man has directed his attention to the problem; and historical record is also absent. The best available evidence is that of paleontology: here the relative positions of the layers of the earth’s crust enable us to deduce their temporal sequence—and naturally, that of the organisms whose fossil remains they embalm—with a great deal of accuracy.[3]
We can scarcely ever observe the direct transition from the forms of life in an older to those in a younger stratum, nor can we absolutely prove their genetic relationship. But in a vast number of cases it is abundantly clear that the later type of organization is descended from the former—that a group of forms in the younger stratum had its origin in one or more species of the group to which the forms in the older stratum belong. Sometimes, however, as in many groups of mammals, the gaps are few and small, the seriation almost complete. In any event we have here evidence which, so far as it goes, is perfectly admissible for the main lines and for many of the smaller branches of evolutionary descent.
Unfortunately, it does not go very far—or, we had better say, it is of restricted application. By the time we find well-preserved fossils in the rocks, the main groups of the animal kingdom and their chief subdivisions had been already differentiated, with the one important exception of the vertebrates; while time, heat, and pressure have so modified the earlier strata as to destroy the fossil forefathers of insects, molluscs, crustacea, and the rest, which they must have contained.
Within the vertebrate stock, then, we can learn a great deal from the semi-direct methods of paleontology: but for the history of the other groups and for their origin and interrelations, we are driven back upon comparative anatomy and embryology, into another field of more circumstantial evidence. When, for instance, we find that the fore-limbs of bat, bird, whale, horse, and man, although so different in function and in detail of structure, are yet built upon the same general plan, and upon a plan wholly different from that of the limbs, say, of a spider or an insect, we must either deny reason and say that this similarity means nothing; or assume that its cause is supernatural, outside the province of science, that it is the expression of some eternal Idea, or some plan of a personal creator (in which case, be it noted, the idea or the plan often appears to our intellect as unreasonable and indeed stupid); or finally that it implies community of origin with later divergence of development. When we are dealing with the smaller sub-divisions of some larger group, this method too gives us information of the same order of accuracy as does paleontology: but when we try to understand the relationships of these larger groups, then we are forced to renounce any claim to detailed knowledge. In broad outline, however, a great deal still remains, and this broad outline we can employ for our valuation of the whole sweep of biological progress, just as we can use the greater accuracy of vertebrate paleontology and comparative morphology to fill in the detail within a restricted field of its operation. From these various evidences, direct and indirect, we can paint for ourselves a picture of the evolution of life which, in spite of inevitable gaps and errors, is in its main features adequate and true.
Let us not be misled by the fact that disputes can and justifiably do arise over details: as Professor Bateson put it recently[4]:—