Going now over to mammals, the first thing which strikes us is the overwhelming numerical predominance of social species over those few carnivores which do not associate. The plateaus, the Alpine tracts, and the Steppes of the Old and New World are stocked with herds of deer, antelopes, gazelles, fallow deer, buffaloes, wild goats and sheep, all of which are sociable animals. When the Europeans came to settle in America, they found it so densely peopled with buffaloes, that pioneers had to stop their advance when a column of migrating buffaloes came to cross the route they followed; the march past of the dense column lasting sometimes for two and three days. And when the Russians took possession of Siberia they found it so densely peopled with deer, antelopes, squirrels, and other sociable animals, that the very conquest of Siberia was nothing but a hunting expedition which lasted for two hundred years; while the grass plains of Eastern Africa are still covered with herds composed of zebra, the hartebeest, and other antelopes.
Not long ago the small streams of Northern America and Northern Siberia were peopled with colonies of beavers, and up to the seventeenth century like colonies swarmed in Northern Russia. The flat lands of the four great continents are still covered with countless colonies of mice, ground-squirrels, marmots, and other rodents. In the lower latitudes of Asia and Africa the forests are still the abode of numerous families of elephants, rhinoceroses, and numberless societies of monkeys. In the far north the reindeer aggregate in numberless herds; while still further north we find the herds of the musk-oxen and numberless bands of polar foxes. The coasts of the ocean are enlivened by flocks of seals and morses; its waters, by shoals of sociable cetaceans; and even in the depths of the great plateau of Central Asia we find herds of wild horses, wild donkeys, wild camels, and wild sheep. All these mammals live in societies and nations sometimes numbering hundreds of thousands of individuals, although now, after three centuries of gunpowder civilization, we find but the debris of the immense aggregations of old. How trifling, in comparison with them, are the numbers of the carnivores! And how false, therefore, is the view of those who speak of the animal world as if nothing were to be seen in it but lions and hyenas plunging their bleeding teeth into the flesh of their victims! One might as well imagine that the whole of human life is nothing but a succession of war massacres.
Association and mutual aid are the rule with mammals. We find social habits even among the carnivores, and we can only name the cat tribe (lions, tigers, leopards, etc.) as a division the members of which decidedly prefer isolation to society, and are but seldom met with even in small groups. And yet, even among lions "this is a very common practice to hunt in company."(10) The two tribes of the civets (Viverridae) and the weasels (Mustelidae) might also be characterized by their isolated life, but it is a fact that during the last century the common weasel was more sociable than it is now; it was seen then in larger groups in Scotland and in the Unterwalden canton of Switzerland. As to the great tribe of the dogs, it is eminently sociable, and association for hunting purposes may be considered as eminently characteristic of its numerous species. It is well known, in fact, that wolves gather in packs for hunting, and Tschudi left an excellent description of how they draw up in a half-circle, surround a cow which is grazing on a mountain slope, and then, suddenly appearing with a loud barking, make it roll in the abyss.(11) Audubon, in the thirties, also saw the Labrador wolves hunting in packs, and one pack following a man to his cabin, and killing the dogs. During severe winters the packs of wolves grow so numerous as to become a danger for human settlements, as was the case in France some five-and-forty years ago. In the Russian Steppes they never attack the horses otherwise than in packs; and yet they have to sustain bitter fights, during which the horses (according to Kohl's testimony) sometimes assume offensive warfare, and in such cases, if the wolves do not retreat promptly, they run the risk of being surrounded by the horses and killed by their hoofs. The prairie-wolves (Canis latrans) are known to associate in bands of from twenty to thirty individuals when they chase a buffalo occasionally separated from its herd.(12) Jackals, which are most courageous and may be considered as one of the most intelligent representatives of the dog tribe, always hunt in packs; thus united, they have no fear of the bigger carnivores.(13) As to the wild dogs of Asia (the Kholzuns, or Dholes), Williamson saw their large packs attacking all larger animals save elephants and rhinoceroses, and overpowering bears and tigers. Hyenas always live in societies and hunt in packs, and the hunting organizations of the painted lycaons are highly praised by Cumming. Nay, even foxes, which, as a rule, live isolated in our civilized countries, have been seen combining for hunting purposes.(14) As to the polar fox, it is—or rather was in Steller's time—one of the most sociable animals; and when one reads Steller's description of the war that was waged by Behring's unfortunate crew against these intelligent small animals, one does not know what to wonder at most: the extraordinary intelligence of the foxes and the mutual aid they displayed in digging out food concealed under cairns, or stored upon a pillar (one fox would climb on its top and throw the food to its comrades beneath), or the cruelty of man, driven to despair by the numerous packs of foxes. Even some bears live in societies where they are not disturbed by man. Thus Steller saw the black bear of Kamtchatka in numerous packs, and the polar bears are occasionally found in small groups. Even the unintelligent insectivores do not always disdain association.
However, it is especially with the rodents, the ungulata, and the ruminants that we find a highly developed practice of mutual aid. The squirrels are individualist to a great extent. Each of them builds its own comfortable nest, and accumulates its own provision. Their inclinations are towards family life, and Brehm found that a family of squirrels is never so happy as when the two broods of the same year can join together with their parents in a remote corner of a forest. And yet they maintain social relations. The inhabitants of the separate nests remain in a close intercourse, and when the pine-cones become rare in the forest they inhabit, they emigrate in bands. As to the black squirrels of the Far West, they are eminently sociable. Apart from the few hours given every day to foraging, they spend their lives in playing in numerous parties. And when they multiply too rapidly in a region, they assemble in bands, almost as numerous as those of locusts, and move southwards, devastating the forests, the fields, and the gardens; while foxes, polecats, falcons, and nocturnal birds of prey follow their thick columns and live upon the individuals remaining behind. The ground-squirrel—a closely-akin genus—is still more sociable. It is given to hoarding, and stores up in its subterranean halls large amounts of edible roots and nuts, usually plundered by man in the autumn. According to some observers, it must know something of the joys of a miser. And yet it remains sociable. It always lives in large villages, and Audubon, who opened some dwellings of the hackee in the winter, found several individuals in the same apartment; they must have stored it with common efforts.
The large tribe, of the marmots, which includes the three large genuses of Arctomys, Cynomys, and Spermophilus, is still more sociable and still more intelligent. They also prefer having each one its own dwelling; but they live in big villages. That terrible enemy of the crops of South Russia—the souslik—of which some ten millions are exterminated every year by man alone, lives in numberless colonies; and while the Russian provincial assemblies gravely discuss the means of getting rid of this enemy of society, it enjoys life in its thousands in the most joyful way. Their play is so charming that no observer could refrain from paying them a tribute of praise, and from mentioning the melodious concerts arising from the sharp whistlings of the males and the melancholic whistlings of the females, before—suddenly returning to his citizen's duties—he begins inventing the most diabolic means for the extermination of the little robbers. All kinds of rapacious birds and beasts of prey having proved powerless, the last word of science in this warfare is the inoculation of cholera! The villages of the prairie-dogs in America are one of the loveliest sights. As far as the eye can embrace the prairie, it sees heaps of earth, and on each of them a prairie-dog stands, engaged in a lively conversation with its neighbours by means of short barkings. As soon as the approach of man is signalled, all plunge in a moment into their dwellings; all have disappeared as by enchantment. But if the danger is over, the little creatures soon reappear. Whole families come out of their galleries and indulge in play. The young ones scratch one another, they worry one another, and display their gracefulness while standing upright, and in the meantime the old ones keep watch. They go visiting one another, and the beaten footpaths which connect all their heaps testify to the frequency of the visitations. In short, the best naturalists have written some of their best pages in describing the associations of the prairie-dogs of America, the marmots of the Old World, and the polar marmots of the Alpine regions. And yet, I must make, as regards the marmots, the same remark as I have made when speaking of the bees. They have maintained their fighting instincts, and these instincts reappear in captivity. But in their big associations, in the face of free Nature, the unsociable instincts have no opportunity to develop, and the general result is peace and harmony.
Even such harsh animals as the rats, which continually fight in our cellars, are sufficiently intelligent not to quarrel when they plunder our larders, but to aid one another in their plundering expeditions and migrations, and even to feed their invalids. As to the beaver-rats or musk-rats of Canada, they are extremely sociable. Audubon could not but admire "their peaceful communities, which require only being left in peace to enjoy happiness." Like all sociable animals, they are lively and playful, they easily combine with other species, and they have attained a very high degree of intellectual development. In their villages, always disposed on the shores of lakes and rivers, they take into account the changing level of water; their domeshaped houses, which are built of beaten clay interwoven with reeds, have separate corners for organic refuse, and their halls are well carpeted at winter time; they are warm, and, nevertheless, well ventilated. As to the beavers, which are endowed, as known, with a most sympathetic character, their astounding dams and villages, in which generations live and die without knowing of any enemies but the otter and man, so wonderfully illustrate what mutual aid can achieve for the security of the species, the development of social habits, and the evolution of intelligence, that they are familiar to all interested in animal life. Let me only remark that with the beavers, the muskrats, and some other rodents, we already find the feature which will also be distinctive of human communities—that is, work in common.
I pass in silence the two large families which include the jerboa, the chinchilla, the biscacha, and the tushkan, or underground hare of South Russia, though all these small rodents might be taken as excellent illustrations of the pleasures derived by animals from social life.(15) Precisely, the pleasures; because it is extremely difficult to say what brings animals together—the needs of mutual protection, or simply the pleasure of feeling surrounded by their congeners. At any rate, our common hares, which do not gather in societies for life in common, and which are not even endowed with intense parental feelings, cannot live without coming together for play. Dietrich de Winckell, who is considered to be among the best acquainted with the habits of hares, describes them as passionate players, becoming so intoxicated by their play that a hare has been known to take an approaching fox for a playmate.(16) As to the rabbit, it lives in societies, and its family life is entirely built upon the image of the old patriarchal family; the young ones being kept in absolute obedience to the father and even the grandfather.(17) And here we have the example of two very closely-allied species which cannot bear each other—not because they live upon nearly the same food, as like cases are too often explained, but most probably because the passionate, eminently-individualist hare cannot make friends with that placid, quiet, and submissive creature, the rabbit. Their tempers are too widely different not to be an obstacle to friendship.
Life in societies is again the rule with the large family of horses, which includes the wild horses and donkeys of Asia, the zebras, the mustangs, the cimarrones of the Pampas, and the half-wild horses of Mongolia and Siberia. They all live in numerous associations made up of many studs, each of which consists of a number of mares under the leadership of a male. These numberless inhabitants of the Old and the New World, badly organized on the whole for resisting both their numerous enemies and the adverse conditions of climate, would soon have disappeared from the surface of the earth were it not for their sociable spirit. When a beast of prey approaches them, several studs unite at once; they repulse the beast and sometimes chase it: and neither the wolf nor the bear, not even the lion, can capture a horse or even a zebra as long as they are not detached from the herd. When a drought is burning the grass in the prairies, they gather in herds of sometimes 10,000 individuals strong, and migrate. And when a snow-storm rages in the Steppes, each stud keeps close together, and repairs to a protected ravine. But if confidence disappears, or the group has been seized by panic, and disperses, the horses perish and the survivors are found after the storm half dying from fatigue. Union is their chief arm in the struggle for life, and man is their chief enemy. Before his increasing numbers the ancestors of our domestic horse (the Equus Przewalskii, so named by Polyakoff) have preferred to retire to the wildest and least accessible plateaus on the outskirts of Thibet, where they continue to live, surrounded by carnivores, under a climate as bad as that of the Arctic regions, but in a region inaccessible to man.(18)
Many striking illustrations of social life could be taken from the life of the reindeer, and especially of that large division of ruminants which might include the roebucks, the fallow deer, the antelopes, the gazelles, the ibex, and, in fact, the whole of the three numerous families of the Antelopides, the Caprides, and the Ovides. Their watchfulness over the safety of their herds against attacks of carnivores; the anxiety displayed by all individuals in a herd of chamois as long as all of them have not cleared a difficult passage over rocky cliffs, the adoption of orphans; the despair of the gazelle whose mate, or even comrade of the same sex, has been killed; the plays of the youngsters, and many other features, could be mentioned. But perhaps the most striking illustration of mutual support is given by the occasional migrations of fallow deer, such as I saw once on the Amur. When I crossed the high plateau and its border ridge, the Great Khingan, on my way from Transbaikalia to Merghen, and further travelled over the high prairies on my way to the Amur, I could ascertain how thinly-peopled with fallow deer these mostly uninhabited regions are.(19) Two years later I was travelling up the Amur, and by the end of October reached the lower end of that picturesque gorge which the Amur pierces in the Dousse-alin (Little Khingan) before it enters the lowlands where it joins the Sungari. I found the Cossacks in the villages of that gorge in the greatest excitement, because thousands and thousands of fallow deer were crossing the Amur where it is narrowest, in order to reach the lowlands. For several days in succession, upon a length of some forty miles up the river, the Cossacks were butchering the deer as they crossed the Amur, in which already floated a good deal of ice. Thousands were killed every day, and the exodus nevertheless continued. Like migrations were never seen either before or since, and this one must have been called for by an early and heavy snow-fall in the Great Khingan, which compelled the deer to make a desperate attempt at reaching the lowlands in the east of the Dousse mountains. Indeed, a few days later the Dousse-alin was also buried under snow two or three feet deep. Now, when one imagines the immense territory (almost as big as Great Britain) from which the scattered groups of deer must have gathered for a migration which was undertaken under the pressure of exceptional circumstances, and realizes the difficulties which had to be overcome before all the deer came to the common idea of crossing the Amur further south, where it is narrowest, one cannot but deeply admire the amount of sociability displayed by these intelligent animals. The fact is not the less striking if we remember that the buffaloes of North America displayed the same powers of combination. One saw them grazing in great numbers in the plains, but these numbers were made up by an infinity of small groups which never mixed together. And yet, when necessity arose, all groups, however scattered over an immense territory, came together and made up those immense columns, numbering hundreds of thousands of individuals, which I mentioned on a preceding page.
I also ought to say a few words at least about the "compound families" of the elephants, their mutual attachment, their deliberate ways in posting sentries, and the feelings of sympathy developed by such a life of close mutual support.(20) I might mention the sociable feelings of those disreputable creatures the wild boars, and find a word of praise for their powers of association in the case of an attack by a beast of prey.(21) The hippopotamus and the rhinoceros, too, would occupy a place in a work devoted to animal sociability. Several striking pages might be given to the sociability and mutual attachment of the seals and the walruses; and finally, one might mention the most excellent feelings existing among the sociable cetaceans. But I have to say yet a few words about the societies of monkeys, which acquire an additional interest from their being the link which will bring us to the societies of primitive men.