The correctness of the above views struck most of the Russian zoologists present, and Syevertsoff, whose work is well known to ornithologists and geographers, supported them and illustrated them by a few more examples. He mentioned sone of the species of falcons which have "an almost ideal organization for robbery," and nevertheless are in decay, while other species of falcons, which practise mutual help, do thrive. "Take, on the other side, a sociable bird, the duck," he said; "it is poorly organized on the whole, but it practises mutual support, and it almost invades the earth, as may be judged from its numberless varieties and species."
The readiness of the Russian zoologists to accept Kessler's views seems quite natural, because nearly all of them have had opportunities of studying the animal world in the wide uninhabited regions of Northern Asia and East Russia; and it is impossible to study like regions without being brought to the same ideas. I recollect myself the impression produced upon me by the animal world of Siberia when I explored the Vitim regions in the company of so accomplished a zoologist as my friend Polyakoff was. We both were under the fresh impression of the Origin of Species, but we vainly looked for the keen competition between animals of the same species which the reading of Darwin's work had prepared us to expect, even after taking into account the remarks of the third chapter (p. 54). We saw plenty of adaptations for struggling, very often in common, against the adverse circumstances of climate, or against various enemies, and Polyakoff wrote many a good page upon the mutual dependency of carnivores, ruminants, and rodents in their geographical distribution; we witnessed numbers of facts of mutual support, especially during the migrations of birds and ruminants; but even in the Amur and Usuri regions, where animal life swarms in abundance, facts of real competition and struggle between higher animals of the same species came very seldom under my notice, though I eagerly searched for them. The same impression appears in the works of most Russian zoologists, and it probably explains why Kessler's ideas were so welcomed by the Russian Darwinists, whilst like ideas are not in vogue amidst the followers of Darwin in Western Europe.
The first thing which strikes us as soon as we begin studying the struggle for existence under both its aspects—direct and metaphorical—is the abundance of facts of mutual aid, not only for rearing progeny, as recognized by most evolutionists, but also for the safety of the individual, and for providing it with the necessary food. With many large divisions of the animal kingdom mutual aid is the rule. Mutual aid is met with even amidst the lowest animals, and we must be prepared to learn some day, from the students of microscopical pond-life, facts of unconscious mutual support, even from the life of micro-organisms. Of course, our knowledge of the life of the invertebrates, save the termites, the ants, and the bees, is extremely limited; and yet, even as regards the lower animals, we may glean a few facts of well-ascertained cooperation. The numberless associations of locusts, vanessae, cicindelae, cicadae, and so on, are practically quite unexplored; but the very fact of their existence indicates that they must be composed on about the same principles as the temporary associations of ants or bees for purposes of migration. As to the beetles, we have quite well-observed facts of mutual help amidst the burying beetles (Necrophorus). They must have some decaying organic matter to lay their eggs in, and thus to provide their larvae with food; but that matter must not decay very rapidly. So they are wont to bury in the ground the corpses of all kinds of small animals which they occasionally find in their rambles. As a rule, they live an isolated life, but when one of them has discovered the corpse of a mouse or of a bird, which it hardly could manage to bury itself, it calls four, six, or ten other beetles to perform the operation with united efforts; if necessary, they transport the corpse to a suitable soft ground; and they bury it in a very considerate way, without quarrelling as to which of them will enjoy the privilege of laying its eggs in the buried corpse. And when Gleditsch attached a dead bird to a cross made out of two sticks, or suspended a toad to a stick planted in the soil, the little beetles would in the same friendly way combine their intelligences to overcome the artifice of Man. The same combination of efforts has been noticed among the dung-beetles.
Even among animals standing at a somewhat lower stage of organization we may find like examples. Some land-crabs of the West Indies and North America combine in large swarms in order to travel to the sea and to deposit therein their spawn; and each such migration implies concert, co-operation, and mutual support. As to the big Molucca crab (Limulus), I was struck (in 1882, at the Brighton Aquarium) with the extent of mutual assistance which these clumsy animals are capable of bestowing upon a comrade in case of need. One of them had fallen upon its back in a corner of the tank, and its heavy saucepan-like carapace prevented it from returning to its natural position, the more so as there was in the corner an iron bar which rendered the task still more difficult. Its comrades came to the rescue, and for one hour's time I watched how they endeavoured to help their fellow-prisoner. They came two at once, pushed their friend from beneath, and after strenuous efforts succeeded in lifting it upright; but then the iron bar would prevent them from achieving the work of rescue, and the crab would again heavily fall upon its back. After many attempts, one of the helpers would go in the depth of the tank and bring two other crabs, which would begin with fresh forces the same pushing and lifting of their helpless comrade. We stayed in the Aquarium for more than two hours, and, when leaving, we again came to cast a glance upon the tank: the work of rescue still continued! Since I saw that, I cannot refuse credit to the observation quoted by Dr. Erasmus Darwin—namely, that "the common crab during the moulting season stations as sentinel an unmoulted or hard-shelled individual to prevent marine enemies from injuring moulted individuals in their unprotected state."(5)
Facts illustrating mutual aid amidst the termites, the ants, and the bees are so well known to the general reader, especially through the works of Romanes, L. Buchner, and Sir John Lubbock, that I may limit my remarks to a very few hints.(6) If we take an ants' nest, we not only see that every description of work-rearing of progeny, foraging, building, rearing of aphides, and so on—is performed according to the principles of voluntary mutual aid; we must also recognize, with Forel, that the chief, the fundamental feature of the life of many species of ants is the fact and the obligation for every ant of sharing its food, already swallowed and partly digested, with every member of the community which may apply for it. Two ants belonging to two different species or to two hostile nests, when they occasionally meet together, will avoid each other. But two ants belonging to the same nest or to the same colony of nests will approach each other, exchange a few movements with the antennae, and "if one of them is hungry or thirsty, and especially if the other has its crop full … it immediately asks for food." The individual thus requested never refuses; it sets apart its mandibles, takes a proper position, and regurgitates a drop of transparent fluid which is licked up by the hungry ant. Regurgitating food for other ants is so prominent a feature in the life of ants (at liberty), and it so constantly recurs both for feeding hungry comrades and for feeding larvae, that Forel considers the digestive tube of the ants as consisting of two different parts, one of which, the posterior, is for the special use of the individual, and the other, the anterior part, is chiefly for the use of the community. If an ant which has its crop full has been selfish enough to refuse feeding a comrade, it will be treated as an enemy, or even worse. If the refusal has been made while its kinsfolk were fighting with some other species, they will fall back upon the greedy individual with greater vehemence than even upon the enemies themselves. And if an ant has not refused to feed another ant belonging to an enemy species, it will be treated by the kinsfolk of the latter as a friend. All this is confirmed by most accurate observation and decisive experiments.(7)
In that immense division of the animal kingdom which embodies more than one thousand species, and is so numerous that the Brazilians pretend that Brazil belongs to the ants, not to men, competition amidst the members of the same nest, or the colony of nests, does not exist. However terrible the wars between different species, and whatever the atrocities committed at war-time, mutual aid within the community, self-devotion grown into a habit, and very often self-sacrifice for the common welfare, are the rule. The ants and termites have renounced the "Hobbesian war," and they are the better for it. Their wonderful nests, their buildings, superior in relative size to those of man; their paved roads and overground vaulted galleries; their spacious halls and granaries; their corn-fields, harvesting and "malting" of grain;(8) their, rational methods of nursing their eggs and larvae, and of building special nests for rearing the aphides whom Linnaeus so picturesquely described as "the cows of the ants"; and, finally, their courage, pluck, and, superior intelligence—all these are the natural outcome of the mutual aid which they practise at every stage of their busy and laborious lives. That mode of life also necessarily resulted in the development of another essential feature of the life of ants: the immense development of individual initiative which, in its turn, evidently led to the development of that high and varied intelligence which cannot but strike the human observer.(9)
If we knew no other facts from animal life than what we know about the ants and the termites, we already might safely conclude that mutual aid (which leads to mutual confidence, the first condition for courage) and individual initiative (the first condition for intellectual progress) are two factors infinitely more important than mutual struggle in the evolution of the animal kingdom. In fact, the ant thrives without having any of the "protective" features which cannot be dispensed with by animals living an isolated life. Its colour renders it conspicuous to its enemies, and the lofty nests of many species are conspicuous in the meadows and forests. It is not protected by a hard carapace, and its stinging apparatus, however dangerous when hundreds of stings are plunged into the flesh of an animal, is not of a great value for individual defence; while the eggs and larvae of the ants are a dainty for a great number of the inhabitants of the forests. And yet the ants, in their thousands, are not much destroyed by the birds, not even by the ant-eaters, and they are dreaded by most stronger insects. When Forel emptied a bagful of ants in a meadow, he saw that "the crickets ran away, abandoning their holes to be sacked by the ants; the grasshoppers and the crickets fled in all directions; the spiders and the beetles abandoned their prey in order not to become prey themselves;" even the nests of the wasps were taken by the ants, after a battle during which many ants perished for the safety of the commonwealth. Even the swiftest insects cannot escape, and Forel often saw butterflies, gnats, flies, and so on, surprised and killed by the ants. Their force is in mutual support and mutual confidence. And if the ant—apart from the still higher developed termites—stands at the very top of the whole class of insects for its intellectual capacities; if its courage is only equalled by the most courageous vertebrates; and if its brain—to use Darwin's words—"is one of the most marvellous atoms of matter in the world, perhaps more so than the brain of man," is it not due to the fact that mutual aid has entirely taken the place of mutual struggle in the communities of ants?
The same is true as regards the bees. These small insects, which so easily might become the prey of so many birds, and whose honey has so many admirers in all classes of animals from the beetle to the bear, also have none of the protective features derived from mimicry or otherwise, without which an isolatedly living insect hardly could escape wholesale destruction; and yet, owing to the mutual aid they practise, they obtain the wide extension which we know and the intelligence we admire, By working in common they multiply their individual forces; by resorting to a temporary division of labour combined with the capacity of each bee to perform every kind of work when required, they attain such a degree of well-being and safety as no isolated animal can ever expect to achieve however strong or well armed it may be. In their combinations they are often more successful than man, when he neglects to take advantage of a well-planned mutual assistance. Thus, when a new swarm of bees is going to leave the hive in search of a new abode, a number of bees will make a preliminary exploration of the neighbourhood, and if they discover a convenient dwelling-place—say, an old basket, or anything of the kind—they will take possession of it, clean it, and guard it, sometimes for a whole week, till the swarm comes to settle therein. But how many human settlers will perish in new countries simply for not having understood the necessity of combining their efforts! By combining their individual intelligences they succeed in coping with adverse circumstances, even quite unforeseen and unusual, like those bees of the Paris Exhibition which fastened with their resinous propolis the shutter to a glass-plate fitted in the wall of their hive. Besides, they display none of the sanguinary proclivities and love of useless fighting with which many writers so readily endow animals. The sentries which guard the entrance to the hive pitilessly put to death the robbing bees which attempt entering the hive; but those stranger bees which come to the hive by mistake are left unmolested, especially if they come laden with pollen, or are young individuals which can easily go astray. There is no more warfare than is strictly required.
The sociability of the bees is the more instructive as predatory instincts and laziness continue to exist among the bees as well, and reappear each time that their growth is favoured by some circumstances. It is well known that there always are a number of bees which prefer a life of robbery to the laborious life of a worker; and that both periods of scarcity and periods of an unusually rich supply of food lead to an increase of the robbing class. When our crops are in and there remains but little to gather in our meadows and fields, robbing bees become of more frequent occurrence; while, on the other side, about the sugar plantations of the West Indies and the sugar refineries of Europe, robbery, laziness, and very often drunkenness become quite usual with the bees. We thus see that anti-social instincts continue to exist amidst the bees as well; but natural selection continually must eliminate them, because in the long run the practice of solidarity proves much more advantageous to the species than the development of individuals endowed with predatory inclinations. The cunningest and the shrewdest are eliminated in favour of those who understand the advantages of sociable life and mutual support.
Certainly, neither the ants, nor the bees, nor even the termites, have risen to the conception of a higher solidarity embodying the whole of the species. In that respect they evidently have not attained a degree of development which we do not find even among our political, scientific, and religious leaders. Their social instincts hardly extend beyond the limits of the hive or the nest. However, colonies of no less than two hundred nests, belonging to two different species (Formica exsecta and F. pressilabris) have been described by Forel on Mount Tendre and Mount Saleve; and Forel maintains that each member of these colonies recognizes every other member of the colony, and that they all take part in common defence; while in Pennsylvania Mr. MacCook saw a whole nation of from 1,600 to 1,700 nests of the mound-making ant, all living in perfect intelligence; and Mr. Bates has described the hillocks of the termites covering large surfaces in the "campos"—some of the nests being the refuge of two or three different species, and most of them being connected by vaulted galleries or arcades.(10) Some steps towards the amalgamation of larger divisions of the species for purposes of mutual protection are thus met with even among the invertebrate animals.