The seed contains a miniature plant, or embryo. The embryo usually has three parts that have received names: the stemlet, or caulicle; the seed-leaf, or cotyledon (usually 1 or 2); the bud, or plumule, lying between or above the cotyledons. These parts are well seen in the common bean (Fig. [15]), particularly when the seed has been soaked for a few hours. One of the large cotyledons—comprising half of the bean—is shown at R. The caulicle is at O. The plumule is shown at A. The cotyledons are attached to the caulicle at F: this point may be taken as the first node or joint.
Fig. 15.— Parts of the Bean.
R, cotyledon; O, caulicle; A, plumule; F, first node.
The Number of Seed-leaves.—All plants having two seed-leaves belong to the group called dicotyledons. Such seeds in many cases split readily in halves, e.g. a bean. Some plants have only one seed-leaf in a seed. They form a group of plants called monocotyledons. Indian corn is an example of a plant with only one seed-leaf: a grain of corn does not split into halves as a bean does. Seeds of the pine family contain more than two cotyledons, but for our purposes they may be associated with the dicotyledons, although really forming a different group.
These two groups—the dicotyledons and the monocotyledons—represent two great natural divisions of the vegetable kingdom. The dicotyledons contain the woody bark-bearing trees and bushes (except conifers), and most of the herbs of temperate climates except the grasses, sedges, rushes, lily tribes, and orchids. The flower-parts are usually in fives or multiples of five, the leaves mostly netted-veined, the bark or rind distinct, and the stem often bearing a pith at the centre. The monocotyledons usually have the flower-parts in threes or multiples of three, the leaves long and parallel-veined, the bark not separable, and the stem without a central pith.
Fig. 16.—External Parts of Bean.
Every seed is provided with food to support the germinating plant. Commonly this food is starch. The food may be stored in the cotyledons, as in bean, pea, squash; or outside the cotyledons, as in castor bean, pine, Indian corn. When the food is outside or around the embryo, it is usually called endosperm.
Seed-coats; Markings on Seed.—The embryo and endosperm are inclosed within a covering made of two or more layers and known as the seed-coats. Over the point of the caulicle is a minute hole or a thin place in the coats known as the micropyle. This is the point at which the pollen-tube entered the forming ovule and through which the caulicle breaks in germination. The micropyle is shown at M in Fig. [16]. The scar where the seed broke from its funiculus (or stalk that attached it to its pod) is named the hilum. It occupies a third of the length of the bean in Fig. [16]. The hilum and micropyle are always present in seeds, but they are not always close together. In many cases it is difficult to identify the micropyle in the dormant seed, but its location is at once shown by the protruding caulicle as germination begins. Opposite the micropyle in the bean (at the other end of the hilum) is an elevation known as the raphe. This is formed by a union of the funiculus, or seed-stalk, with the seed-coats, and through it food was transferred for the development of the seed, but it is now functionless.