When the Medusæ travel, their convex part is always kept in advance, and slightly oblique. If they are touched while swimming, even lightly, they contract their tentacula, fold up their umbrella, and sink into the sea. Like Ehrenberg, M. Kölliker thought he discovered visual and auditory organs in an Oceania, and Gegenbauer thought he detected them in other genera, such as Rhizostoma and Pelagia. The eyes are said to consist of certain small, hemispherical, cellulose, coloured masses, in which are sunk small crystalline globules, the free parts of which are perfectly naked. The supposed auditory apparatus is seated close to these organs; they are small vesicles filled with liquid; the eyes having neither pupil nor cornea, and the ears without opening or arch.
But it is in their reproduction that these evanescent beings present the most marvellous phenomena. At one period of the year the Medusæ are charged with numbers of very minute eggs, of the most lively colours, which are suspended in large festoons from their floating bodies. In some cases these eggs develop themselves grafted to their bodies, and are only detached at maturity. In other cases the larvæ produced bear no resemblance to the mother; they are elongated and vermiform, broad at their extremity; we speak of the microscopic leeches, which have vibrating cilia, scarcely perceptible, by which they execute the most lively motions. At the end of a certain time they are transformed into polyps, and furnished with eight tentacula. This preparatory sort of animal seems to possess the faculty of reproduction by means of certain buds or tubercles which develop themselves on the surface of the body, and also by filaments which start up here and there, so that a single individual originates a numerous colony. This polyp is subjected to a transformation still more remarkable; its structure becomes complex, its body articulate, and it seems to be composed of a dozen disks piled one upon the other, like the jars of a voltaic pile; the upper disk is convex, and is separated from the colony after a convulsive effort; it becomes free, and an excessively small, star-like Medusa is the result; every disk, that is, every individual, is isolated one after the other in the same manner.
Thus of the sexual zoophytes which propagate their kind according to the usual laws; but others engender young which have no resemblance to the parent zoophyte at all: in this respect they are neuter, that is, non-sexual, or agamous. These are produced by budding, or fissiparity, from individuals like themselves. They can also give sexual distinctions; but before this change takes place the creature, which was simple, is transformed into a composite animal, and it is from its disaggregation that individuals having sexual organs are produced, the process being that which has been called alternate generation. It goes on in a perfectly regular manner, although it is a fact that the young never resemble their mothers, but their grandmothers.
This great family of Zoophytes Gosse divides into—
Discophora, having the body in the form of a circular disk, more or less convex and umbrella-shaped, moving by alternate contractions and expansions of the disk.
Fig. 84. æquerea violacea, natural size (Milne Edwards).
Ctenophora, body cylindrical, moving by means of many parallel rims of cilia set in longitudinal lines on the surface.
Sophonophora, body irregular, without central digestive cavity like the others, having sucking organs, and moving by means of a contractile cavity, or by air-vessels.