Included in the Scyphozoa are the large jelly-fish found floating on the sea or cast up on the beach on the British shores.

The Anthozoa include the Sea-anemones, nearly all the Stony Corals, the Sea-fans, the Black Corals, the Dead-men's fingers (Alcyonium), the Sea-pens, and the Precious Coral of commerce.

CLASS I. HYDROZOA

In this Class of Coelenterata two types of body-form may be found. In such a genus as Obelia there is a fixed branching colony of zooids, and each zooid consists of a simple tubular body-wall composed of the two layers of cells, the ectoderm and the endoderm (Fig. 125), terminating distally in a conical mound—the "hypostome"—which is perforated by the mouth and surrounded by a crown of tentacles. This fixed colony, the "hydrosome," feeds and increases in size by gemmation, but does not produce sexual cells. The hydrosome produces at a certain season of the year a number of buds, which develop into small bell-like jelly-fish called the "Medusae," which swim away from the parent stock and produce the sexual cells. The Medusa (Fig. 126) consists of a delicate dome-shaped contractile bell, perforated by radial canals and fringed with tentacles; and from its centre there depends, like the clapper of a bell, a tubular process, the manubrium, which bears the mouth at its extremity. This free-swimming sexual stage in the life-history of Obelia is called the "medusome."

It is difficult to determine whether, in the evolution of the Hydrozoa, the hydrosome preceded the medusome or vice versâ. By some authors the medusome is regarded as a specially modified sexual individual of the hydrosome colony. By others the medusome is regarded as the typical adult Hydrozoon form, and the zooids of the hydrosome as nutritive individuals arrested in their development to give support to it. Whatever may be the right interpretation of the facts, however, it is found that in some forms the medusome stage is more or less degenerate and the hydrosome is predominant, whereas in others the hydrosome is degenerate or inconspicuous and the medusome is predominant. Finally, in some cases there are no traces, even in development, of a medusome stage, and the life-history is completed in the hydrosome, while in others the hydrosome stages are lost and the life-history is completed in the medusome.

If a conspicuous hydrosome stage is represented by H, a conspicuous medusome stage by M, an inconspicuous or degenerate hydrosome stage by h, an inconspicuous or degenerate medusome stage by m, and the fertilised ovum by O, the life-histories of the Hydrozoa may be represented by the following formulæ:—

1.O — H — O(Hydra)
2.O — H — m — O(Sertularia)
3.O — H — M — O(Obelia)
4.O — h — M — O(Liriope)
5.O — M — O(Geryonia)

The structure of the hydrosome is usually very simple. It consists of a branched tube opening by mouths at the ends of the branches and closed at the base. The body-wall is built up of ectoderm and endoderm. Between these layers there is a thin non-cellular lamella, the mesogloea.

In a great many Hydrozoa the ectoderm secretes a chitinous protective tube called the "perisarc." The mouth is usually a small round aperture situated on the summit of the hypostome, and at the base of the hypostome there may be one or two crowns of tentacles or an area bearing irregularly scattered tentacles. The tentacles may be hollow, containing a cavity continuous with the coelenteric cavity of the body; or solid, the endoderm cells arranged in a single row forming an axial support for the ectoderm. The ectoderm of the tentacles is provided with numerous nematocysts, usually arranged in groups or clusters on the distal two-thirds of their length, but sometimes confined to a cap-like swelling at the extremity (capitate tentacles). The hydrosome may be a single zooid producing others asexually by gemmation (or more rarely by fission), which become free from the parent, or it may be a colony of zooids in organic connexion with one another formed by the continuous gemmation of the original zooid derived from the fertilised ovum and its asexually produced offspring. When the hydrosome is a colony of zooids, specialisation of certain individuals for particular functions may occur, and the colony becomes dimorphic or polymorphic.