Stigmaria and its rootlets belong equally to the two plants, and hitherto it has been impossible to tell whether any given specimen of Stigmaria had belonged to a Lepidodendron or a Sigillaria. Between the two genera there certainly existed the closest affinity and similarity in general appearance.
These two genera represent the climax of development of the Lycopod family. In the Lower Mesozoic some large forms are still found, but all through the Mesozoic periods the group dwindled, and in the Tertiary little is known of it, and it seems to have taken the retiring position it occupies to-day.
CHAPTER XV
PAST HISTORIES OF PLANT FAMILIES
VIII. The Horsetails
The horsetails of to-day all belong to the one genus, Equisetum, among the different species of which there is a remarkably close similarity. Most of the species love swampy land, and even grow standing up through water; but some live on the dry clay of ploughed fields. Wherever they grow they usually congregate in large numbers, and form little groves together. They are easily recognized by their delicate stems, branching in bottle-brush fashion, and the small leaves arranged round them in whorls, with their narrow teeth joined to a ring at the base. At the end of some of the branches come the cones, with compactly arranged and simple sporophylls all of one kind. In England most plants of this family are but a few inches or a foot in height, though one species sometimes reaches 6 ft., while in South America there are groves of delicate-stemmed plants 20 ft. high.
The ribbed stems and the whorls of small, finely toothed leaves are the most important external characteristics of the plants, while in their internal anatomy the hollow stems have very little wood, which is arranged in a series of small bundles, each associated with a hollow canal in the ground tissue.
The family stands apart from all others, and even between it and the group of Lycopods there seems to be a big gap across which stretch no bonds of affinity. Has the group always been in a similar position, and stood isolated in a backwater of the stream of plant life?
Fig. 103.—Impression of Leaf Whorl of Equisetites from the Mesozoic Rocks, showing the narrow toothed form of the leaves. (Photo.)
In the late Tertiary period they seem to have held much the same position as they do now, and we learn nothing new of them from rocks of that age. When, however, we come to the Mesozoic, the members of the family are of greater size, though they appear (to judge from their external appearance) to have been practically identical with those now living in all their arrangements. In some beds their impressions are very numerous, but unfortunately most are without any indication of internal structure. Fossils from the Mesozoic are called Equisetites, a name which indicates that they come very close to the living ones in their characters. In the Lower Mesozoic some of these stems seem to have reached the great size of a couple of feet in circumference, but to have no essential difference from the others of the group.
When, however, we come to the Palæozoic rocks we find many specimens with their structure preserved, and we are at once in a very different position as regards the family.