The Rotaline series of the Globigerina family is one of the most numerous and varied of the whole class of Foraminifera; but varied as their forms are, they all bear the characteristic marks which distinguish their order, with this essential difference, that in the genus Globigerina each chamber of the spire has a communication with the central vestibule by a crescent-shaped aperture, while in the Rotalinæ each chamber only communicates by a crescentic aperture with that which precedes and follows it.
In the Rotaline group the internal organization rises successively from the simple porous partition between the chambers, to the double partition with the radiating passages, and from the latter to the double partitions, intermediate skeleton, and complicated system of canals. To these changes the structure of the compound animal necessarily corresponds, for it may be presumed that not only the chambers but all the passages and canals in the interior of the shell are either permanently or occasionally filled with its sarcode body.
However, it is in the Nummuline family that the Foraminifera attain the highest organization of which they are capable. This family surpasses all the Vitreous tribe in the density and toughness of the shell, the fineness of its tubuli, and in the high organization of its canal system. Their forms vary from that resembling a nautilus or ammonite to a flat spiral or cyclical disk, like an Orbitolite, though vastly superior to it in organization both with regard to the animal and to the structure of the shell.
All the species of the genus Nummulite are spiral; in the typical form the last turn of the spire not only completely embraces, but entirely conceals, all that precede it. In general, the form is that of a double convex lens of more or less thickness; some are flat, lenticular, and thinned away to an acute edge, while others may be spheroidal with a round, or obtuse edge. They owe their name to their resemblance to coins, being, in general, nearly circular. Their diameters range from 1⁄16th of an inch to 41⁄2 inches, so that they are the giants of their race; but the most common species vary from 1⁄2 an inch to 1 inch in diameter.
Fig. 101. Section of Faujasina.
[Fig. 101] represents a section of the Nummulite Faujasina near and parallel to the base of the shell. It shows a series of chambers arranged in a flat spiral, and increasing in size from the centre to the last turn of the spire, which embraces and conceals all that precede it. Every segment of the animal is enclosed in a shell of its own, so that they are separated from one another by a double wall and space between; however, they are connected in the spiral direction by narrow passages in the walls.
The segments of the animal in the exterior whorl have direct communication with the water by means of a shelly marginal cord, a, [fig. 101], perforated by multitudes of minute tubes, less than the 1⁄10000 of an inch in diameter, through which threads of sarcode finer than those of a spider’s web can be protruded. These tubuli are so very fine and numerous, that they characterize the Nummuline family.
Fig. 102. Interior of the Operculina.