Changes of form very generally, but not always, are accompanied with a change in regularity: thus a flower habitually bi-lateral may assume the characters of radiating symmetry and vice versâ. Increase or decrease of size very frequently also are co-existent with an alteration in the usual form.
In the case of the arrangement of organs it is often difficult or impossible, in the present state of our knowledge, to determine whether a given arrangement is congenital or acquired subsequently to the first development, whether for instance an isolation of parts be due to primordial separation or to a subsequent disunion of originally combined organs, [see p. 58]. With reference to the changes in the form of organs, however, it is in general more easy to ascertain the proximate cause of the appearance, and thus teratological changes of form may be grouped according as they are due to, 1, arrest of development; 2, undue or excessive development; 3, perverted development; and 4, irregular development; hence the use of the following terms—Stasimorphy, Pleiomorphy, Metamorphy, and Heteromorphy—to include teratological changes really or apparently due to one or other of the causes above mentioned. The classification here adopted is of course to a considerable extent an arbitrary one and subject to correction or modification, as the knowledge of the development of the flowers in the various genera of plants advances.
FOOTNOTES:
[217] The word symmetry has been used in very different senses by different botanists, sometimes as synonymous with "regularity," at other times to express the assumed typical form of a flower. Payer understands it to be that arrangement of parts which permits of the whole flower being divided vertically into two symmetrical halves (bi-lateral symmetry). Others, again, have applied the term symmetry to the number of the parts of the flower, reserving the terms "regularity" or "irregularity" for the form. It is here used in a general sense to express the plan of the flower, and thus includes the arrangement, form, and number of its component elements.
[218] See Baillon, 'Adansonia,' v, 176.
PART I.
STASIMORPHY.[219]
Deviations from the ordinary form of organs arising from stasis or arrest of development are included under this heading.
There are many cases in which the forms proper to a juvenile condition of the plant are retained for a much longer period than ordinary, or even throughout the life of the individual growth goes on, but "development" is checked. Such conditions may even be propagated by seed or bud. It is a very general thing for botanists to consider these cases as reversions to a simpler, primitive type, and this may be so; but on the other hand, they may be degenerations from a complex type, or they may have no direct relation to any antecedent condition. Stasimorphic changes affecting principally the relative size of organs—such, for instance, as the non-development of internodes, or the atrophy or suppression of parts will be found mentioned in the sections relating to those subjects. In the present part those alterations which affect the form of organs principally are treated of.