Nor can we leave the discussion of what plants are without some mention of the thing that really makes up their structure, whether it be a microscopic bacterial organism or the Big Tree of California. For the unit of all animal and plant life is the cell. In its simplest form it is merely a minute sac with a definite wall and inside the wall is a substance known as protoplasm, literally protos, first, and plasma, thing formed. It is protoplasm that forms the living tissue of all plants and animals; it is life itself. No one has ever succeeded in making any, notwithstanding that many learned men have tried for years. Its inclosure in the cell wall, its power of self-division and consequent multiplication of the units, make up those first things about which most of us can never know much, but the end of which we recognize in the beauty of plant life all about us. For only under the highest powers of the microscope may cells be seen and studied. Just as bankers reckon mills as a definite unit of a cent, and yet none of them has ever seen a mill, so we must think of cells as the definite unit of all living things, although most of us will never see a cell. But, unlike the mill, cells may be seen by those equipped to see them, and this study, the development and grouping of them to form all the varied objects that inhabit the plant world, is known as histology. It is literally the internal history of plants and animals, and lies quite outside the scope of this book. What we must never forget is that whatever knowledge we have gained, either from the foregoing account of what plants are, or from our observation of them, is, after all, only a partial notion of them, as unsatisfactory as our estimate of what people really are, from merely looking at the outside of the houses in which they live. The outer form we may know and admire, the inner substance must ever remain for most of us a secret treasure house the value of which is certain, but the key to which we do not possess.
CHAPTER II
PLANT BEHAVIOR
1. Light and Its Importance To the Plant
PRACTICALLY all that has been said in the first chapter relates to what plants are, their organs, or what we may call the architecture or plan of their framework. But what they do with this elaborate structure is as important as what we do with a house that may contain every modern improvement but is never a home until these things have been put to use. One of the chief concerns of any architect is to see to it that the house has as much sunshine by day and as attractive illumination by night as possible. Nature, that greatest of all architects, also sees to it that plants get the utmost necessary sunlight, but for a much more important reason than the mere attractiveness of sunshine, be that ever so beautiful. For light, the life-giver of all green things, is so absolutely essential to plants that experiments to grow them in the dark have always failed, and many gardeners now use electric light in greenhouses in order to prolong the short daylight of winter. It is the lack of light that makes celery blanch.
Plants grown in the house inevitably turn toward the windows, even plants growing against a wall turn their leaves away from it—nowhere can one find living green things that do not find the light as surely and persistently as men try to get their food or their mates. Many examples of this could be given and must have been noticed by everyone.
Sometimes seeds germinate under a barn floor for instance, and the puny pale little plantling reaches out slender stems, all of which turn, as a compass turns to the north, to perhaps a crack of light in one corner of the building. We have already seen how the search for light will carry the slender rattan palms of India hundreds of feet to the topmost leaves of the forest. Individual plants, and, as we shall see later, whole forests make desperate efforts to get to the light. We know already, that the struggle for light is just as bitter as the struggle for food by roots. And finally if, as we have many times proved by experiments, plants die when grown in a dark room, what is it that light does for plants and how is a process carried on that everything leads us to think is of the greatest possible importance? Quite obviously it is not the mere beauty of sunshine dancing upon the landscape, as entrancing a picture as that may be any summer afternoon, with the play of sunshine and shadow on the tracery of foliage. That green color of the foliage, the almost universal green of so much of the earth’s vegetation, restful to tired eyes, providing us with the most pleasant shade, has wrapped up within it the secret of just what sunshine does for plants. For under the magic of light acting upon this greenery one of the most important industries in the world, the manufacture of food, is constantly going on.
LEAVES AS FACTORIES FOR THE MAKING OF FOOD
It must be clear enough from the start that to call a leaf a factory for the making of food forces us to decide at once whether this is a mere way of speaking, or whether, incredible as it may seem, anything as thin as a leaf can really produce food. As we eat lettuce, and millions of cattle graze every day, leaves as food producers win handily on that score. But to understand how food is produced in such a tiny factory demands that we walk about in it for a bit, study the inside of it and especially its many small chambers within which is not only the machinery, but some of the finished product stored up for later use.
Unlike modern factories there are many entrances, from any one of which we can begin our tour of inspection. On the under side of nearly all leaves and on the upper side of some there are scores or even hundreds of small pores called stoma, so small that only with a microscope can they be seen. These entrances through the factory wall, are carefully guarded by a pair of watchmen whose business it is to see neither too much dry air gets in nor too much of the product of the factory gets out. They see to it, also, that waste products are thrown out at the proper time. These watchmen, or guard cells, as they are called, are constantly on the job, work almost automatically, but their chief function is connected with the proper ventilation of the place, and will be discussed later under “How Plants Breathe.”
Once past the entrance it is obvious that we are in one of the strangest of all factories, for none of the rooms are truly square or oblong and their irregularity as to outline would drive your average foreman into profanity. Yet they are certainly divided into distinct classes, at least as to size and as to what the rooms contain. Some are apparently filled with nothing but air and have direct connection through the stoma with the outdoors. These are called intercellular spaces. Others, and these are most important, are filled mostly with the green coloring matter that gives the leaf its color. This substance is known as chlorophyll, its individual units as chloroplasts, or literally, chlorophyll bodies. Quite independently of these chlorophyll cells or rooms, or the intercellular spaces which correspond to halls, there are some large and many small tubes. These are the veins of the leaf and their finer branches and by their direct connection through the stem to the roots, serve as the ducts through which some of the raw materials are brought into the factory.