Remarks.—Merriam (1897a:103) named this jumping mouse as a full species, mentioning that the skull is similar in size and characters to that of Zapus hudsonius but that externally these animals differed in coloration and length of the tail. Hall (1934:377) treated Z. tenellus as a subspecies of Z. hudsonius. He observed that the difference between Z. tenellus, Z. h. alascensis, and Z. h. hudsonius was of the same degree, and, even though intergrading material was not known to him, he considered tenellus only subspecifically distinct from Z. hudsonius. Hall (loc. cit.) tentatively referred to Z. h. tenellus specimens from Indianpoint Lake, 15 mi. NE Barkerville, Cottonwood P. O., and Hazelton, British Columbia. I have seen and compared with the type of Z. tenellus all specimens examined by Hall and agree with him that they are best referred to Z. h. tenellus. Since 1934, several additional localities in British Columbia have yielded specimens. Those from Minaker River and Fort St. John are intermediate in dorsal coloration and in size and shape of the auditory bullae between Zapus hudsonius hudsonius and Z. h. tenellus but in all other characters are most nearly like Z. h. tenellus to which they are here assigned. These intergrades constitute additional evidence that Z. tenellus and Z. hudsonius are only subspecies of a single species.

Specimens examined.—Total, 17, all from British Columbia, distributed as follows: east side Minaker River, 1 mi. W Trutch, 1; Hazelton, 959 ft., 2 (MVZ); 5 mi. W and 3 mi. N Fort St. John, 1; Indianpoint Lake, 15 mi. NE Barkerville, 5 (MVZ); Cottonwood P. O., 3 (MVZ); S end Swan Lake, Vernon, 1200 ft., 2 (MVZ); Kamloops, 3 (USBS).

Marginal records.—British Columbia.—E side Minaker River, 1 mi. W Trutch; 5 mi. W and 3 mi. N Fort St. John; S end Swan Lake, Vernon, 1200 ft.; Kamloops; Hazelton, 959 ft.

Table 5.—Cranial Measurements (in Millimeters) of Zapus.

Number examined, ♂♂ plus ♀♀Breadth of braincaseCondylobasal lengthInterorbital breadthMastoidal breadthLength of maxillary tooth-rowOccipitonasal lengthPalatal lengthZygomatic breadthZygomatic length

Zapus trinotatus eureka, Big Lagoon, California.
13 mean10.321.44.210.93.924.510.412.59.5
max10.622.14.411.14.025.511.513.210.0
min10.020.73.910.03.724.010.512.19.0

Zapus trinotatus montanus, Crater Lake, Oregon.
10 mean10.320.74.510.64.024.310.512.19.5
max10.521.54.711.14.124.810.812.69.8
min10.220.34.210.23.723.510.111.89.0

Lost Cr. R. S., 10 mi. SE McKenzie Bridge, Oregon.
5 mean10.320.94.510.73.924.210.412.19.0
max10.521.34.610.94.124.610.712.29.0
min10.220.44.410.53.823.510.012.09.1

Zapus trinotatus orarius, 3 mi. W Inverness, 800 ft., California.
12 mean9.920.43.810.83.722.610.012.19.3
max10.220.94.111.03.823.510.612.59.8
min9.619.93.710.43.621.79.811.98.5

Zapus trinotatus trinotatus, Old Fort Clatsop, 100 ft., Oregon.
8 mean10.321.14.410.73.824.310.612.39.0
max10.621.84.811.03.925.011.012.710.0
min10.020.24.010.43.723.610.011.99.2

Cayuse Meadow, 3800 ft., 31/2 mi. SW Steamboat Mt’n, Wash.
10 mean10.421.54.511.13.924.610.712.69.7
max10.622.24.811.44.125.411.112.810.0
min10.220.63.810.83.823.710.212.29.3

Snoqualmie Pass, Washington.
5 mean10.621.64.611.14.024.810.912.89.0
max10.722.24.811.54.225.611.213.210.0
min10.421.24.410.93.824.010.512.59.5

Alta Lake, 2200 ft., British Columbia.
5 mean10.621.64.311.34.124.710.812.79.0
max10.921.94.611.54.325.010.913.19.0
min10.521.24.211.03.924.410.712.39.6

Zapus princeps cinereus, Raft River Mt’s, Utah.
4 mean10.521.55.011.14.124.510.912.69.0
max11.022.55.111.44.325.011.612.910.0
min10.321.04.610.73.924.010.412.39.4

Mt. Harrison, 10 mi. S Albion, Idaho.
13 mean10.521.44.810.94.124.910.912.410.1
max10.722.05.111.44.225.511.412.810.8
min10.220.94.510.53.924.310.211.79.8

Zapus princeps curtatus, Pine Forest Mt’s, Nevada.
11 mean10.521.14.710.74.224.610.912.29.7
max10.621.85.011.04.425.311.412.510.0
min10.220.54.210.53.924.010.511.99.4

Zapus princeps idahoensis, several localities near Cody, Wyoming.
24 mean10.521.44.611.04.024.710.912.69.6
max11.222.65.211.44.225.611.513.310.2
min9.920.64.310.53.924.110.212.09.3

5 mi. E Warm Lake, 7000 ft., Idaho.
4 mean10.221.24.311.04.124.410.712.39.0
max10.321.84.411.14.224.811.012.69.0
min10.020.74.210.84.023.910.312.09.1

Summit Smith Mt’n, 7500 ft., Idaho.
9 mean10.221.34.410.94.124.510.712.19.0
max10.522.54.611.34.325.211.212.610.0
min10.020.84.010.53.923.910.311.89.1

2 mi. NE Cooke, 8500 ft., Montana.
6 mean10.421.14.410.94.024.310.512.59.0
max10.621.84.511.54.125.411.112.910.0
min10.220.34.310.73.823.59.912.09.0

Birch Cr., 18 mi. NE Dillon, 7100 ft., Montana.
11 mean10.321.34.311.04.024.510.812.69.6
max10.622.24.611.64.325.511.513.09.9
min9.720.74.010.63.823.710.312.49.2

Zapus princeps idahoensis, Waterton Lakes Park, Alberta.
5 mean10.621.44.610.74.124.510.812.29.0
max10.922.14.711.04.225.211.012.310.0
min10.321.04.410.44.024.210.611.89.3

Zapus princeps kootenayensis, near Rossland, British Columbia.
10 mean10.220.54.410.64.023.710.411.99.2
max10.821.04.810.94.224.410.712.29.5
min9.720.04.110.03.823.29.911.49.0

Zapus princeps luteus, White Mt’s, Arizona.
20 mean10.120.34.910.73.923.810.411.99.7
max10.421.15.111.24.024.810.912.610.2
min9.619.14.310.23.622.59.511.18.9

Espanola, 5000 ft., New Mexico.
3 mean9.819.84.710.53.723.49.911.59.0
max10.020.14.810.63.823.710.311.69.0
min9.719.54.510.33.622.99.611.49.4

Zapus princeps minor, 2 mi. W Fort Totten, 1400 ft., No. Dakota.
11 mean9.920.54.510.63.723.710.511.89.6
max10.120.84.810.83.824.210.712.19.9
min9.720.04.410.43.623.410.211.49.3

Near Bottineau, North Dakota.
4 mean10.120.94.610.63.824.110.812.310.0
max10.221.34.710.93.824.510.912.510.0
min10.120.64.510.43.723.810.712.19.9

Head Eagle Cr., Bear Paw Mt’s, Montana.
7 mean10.020.84.610.73.824.210.712.19.0
max10.521.34.710.94.024.711.112.510.0
min9.820.34.410.53.623.210.311.99.7

N Maple Cr., Cypress Hills, Saskatchewan.
10 mean10.121.24.610.73.924.510.912.39.8
max10.421.74.910.84.024.811.312.710.0
min9.920.44.410.53.624.010.511.89.6

Zapus princeps oregonus, Parker Cr., Warner Mt’s, 5500 ft., Cal.
12 mean10.621.64.711.14.225.011.112.610.2
max11.022.24.911.64.425.711.413.010.9
min10.221.24.310.84.024.510.712.49.9

Cobb Cr., 6 mi. SW Mt’n City, Nevada.
12 mean10.721.65.011.24.125.011.212.610.0
max11.122.15.211.44.325.711.813.010.3
min10.521.04.610.73.824.410.912.29.5

Wisconsin Cr., 8000 ft., Nevada.
10 mean10.621.64.911.24.024.811.112.49.5
max10.822.25.011.54.225.211.412.89.6
min10.321.24.610.83.924.110.612.29.3

North Fork Malheur River, 21 mi. SE Prairie City, 5000 ft., Ore.
10 mean10.621.54.711.34.224.811.012.79.9
max11.222.35.211.64.426.211.713.210.9
min10.020.84.310.94.023.510.512.49.7

Zapus princeps pacificus, North Fork Coffee Cr., 4500 ft., Calif.
8 mean10.421.54.710.93.924.811.112.510.0
max10.822.45.011.44.025.211.413.210.0
min10.020.74.510.73.824.010.512.09.6

Jackson Lake, 5900 ft., California.
7 mean10.421.54.511.14.024.911.012.610.0
max10.622.14.611.54.125.511.412.910.0
min10.120.74.310.53.823.510.012.29.6

Head of Lyle Canyon, 9700 ft., California.
7 mean10.320.84.710.63.824.010.512.39.0
max10.521.85.011.04.024.610.812.710.0
min10.020.04.510.23.523.010.011.89.0

Zapus princeps princeps, Florida, Colorado.
7 mean10.221.44.610.53.724.911.112.39.0
max10.522.34.711.33.825.411.412.510.0
min9.720.74.39.83.523.910.911.99.3

Zapus princeps princeps, Half Way, Colorado.
6 mean10.121.74.610.83.924.911.012.39.0
max10.322.04.811.14.025.811.312.710.0
min10.021.24.510.53.824.210.712.19.6

8 mi. N, 191/2 mi. E Savery, Wyoming.
11 mean10.221.24.510.93.924.510.812.29.7
max10.521.84.711.14.125.011.112.510.2
min10.020.84.210.63.723.710.512.09.3
211/2 mi. S, 241/2 mi. W Douglas, 7600 ft., Wyoming.
11 mean10.121.24.610.94.024.610.812.39.8
max10.421.94.911.24.125.011.212.810.1
min9.920.74.510.83.824.210.512.09.5

Medicine Wheel Ranch, 28 mi. E Lovell, 9000 ft., Wyoming.
20 mean10.321.54.711.24.024.711.012.610.0
max10.622.24.911.54.225.311.412.910.4
min10.020.74.410.83.824.110.612.29.8

Zapus princeps saltator, Stikine River at Glenora, British Columbia.
17 mean10.421.34.410.94.124.310.712.59.6
max10.722.24.511.44.525.011.413.010.0
min9.820.54.110.63.823.310.312.09.3

Hazelton, 959 ft., British Columbia.
15 mean10.421.64.511.04.024.610.912.59.8
max10.822.34.711.64.225.511.412.910.0
min9.920.74.210.73.823.710.511.79.4

Zapus princeps utahensis, near Robertson, 8700 ft., Wyoming.
15 mean10.722.05.011.24.125.411.113.29.9
max11.122.65.111.64.226.411.714.010.3
min10.321.04.710.83.924.610.812.49.6

3 mi. N and 11 mi. E Alpine, 5650 ft., Wyoming.
17 mean10.622.14.711.34.125.311.313.09.9
max11.023.04.911.74.326.211.813.510.5
min10.321.34.4 10.84.024.210.712.19.5

Zapus princeps utahensis, Salamander Lake and Lambs Canyon, 9000 ft., Utah.
9 mean10.922.04.811.24.125.211.113.110.0
max11.322.45.011.34.325.911.413.310.0
min10.721.54.511.03.924.610.712.69.7

Zapus hudsonius acadicus, vicinity of St. Andrews, New Brunswick.
4 mean9.619.84.110.23.523.010.110.89.0
max9.819.94.210.33.723.110.211.19.0
min9.219.73.910.03.322.89.910.49.2

Sebec Lake, Maine.
3 mean9.619.54.310.03.523.010.010.79.0
max9.819.84.510.13.623.310.111.39.0
min9.419.04.09.73.422.69.99.99.0

2 mi. S Center Ossipee, New Hampshire.
10 mean9.619.84.210.13.523.310.010.89.3
max10.020.54.610.43.624.010.311.09.9
min9.218.93.89.73.222.39.610.68.8

Berlin, New York.
6 mean9.519.54.310.13.522.99.810.89.0
max9.920.64.410.63.723.810.611.39.0
min9.218.84.29.53.421.89.310.48.6

Lake Kedgemakooge, Nova Scotia.
5 mean9.519.94.210.33.523.410.211.39.0
max9.720.14.310.43.523.510.411.49.0
min9.319.74.010.33.423.39.911.09.3

Zapus hudsonius alascensis, Lake Clark, Alaska.
2 mean9.519.34.310.03.622.89.910.79.0
max9.619.64.410.03.623.09.910.79.0
min9.419.14.29.93.522.69.910.79.2

Frosty Peak, Yakutat Bay, Alaska.
3 mean9.819.74.210.33.623.510.110.89.0
max10.020.04.510.63.724.210.411.19.0
min9.619.04.09.83.522.59.610.29.2

Zapus hudsonius alascensis, 7 mi. SSE Haines, 10 ft., Alaska.
14 mean9.920.04.210.43.623.710.211.09.4
max10.120.54.410.83.724.610.711.49.8
min9.819.54.010.23.423.09.810.49.0

SW end Dezadeash Lake, Yukon.
2 mean10.019.84.510.53.623.510.211.39.0
max10.120.14.510.53.623.810.411.39.0
min9.819.54.410.53.523.210.011.29.5

Zapus hudsonius americanus, Boyne Falls, Michigan.
8 mean9.518.74.19.73.322.09.511.09.0
max9.819.44.310.03.423.210.011.49.0
min9.318.33.89.43.021.59.010.39.0

Ann Arbor, Michigan.
3 mean9.518.64.29.93.322.09.510.99.0
max9.618.84.410.03.322.49.811.09.0
min9.418.53.89.83.221.99.310.88.9

Montauk Point, L. I., New York.
2 mean9.418.84.39.23.522.29.710.68.0
max9.619.14.49.23.522.59.810.78.0
min9.218.44.29.23.421.99.510.58.9

Mays Landing, New Jersey.
2 mean9.418.84.49.83.422.19.610.98.0
max9.518.84.49.83.522.29.711.08.0
min9.318.74.49.83.222.09.510.88.2

Laurel, Maryland.
3 mean9.118.64.39.63.322.09.510.68.0
max9.318.94.59.73.322.09.710.88.0
min8.918.24.19.53.321.99.210.48.6

Hampton, Virginia.
3 mean9.018.84.19.73.321.99.410.69.0
max9.318.94.19.83.322.09.611.19.0
min8.618.54.09.63.221.89.210.08.9

Zapus hudsonius americanus, Raleigh, North Carolina.
3 mean9.218.84.09.93.422.49.610.98.0
max9.619.74.210.43.523.09.910.99.0
min8.817.93.99.43.221.89.410.88.5

Zapus hudsonius campestris, 3 mi. NW Sundance, 5900 ft., Wyo.
19 mean9.719.94.310.43.623.210.011.19.5
max10.020.84.510.93.824.210.511.810.0
min9.419.23.810.13.422.49.510.79.2

Palmer Gulch, Black Hills, South Dakota.
11 mean9.820.24.310.53.723.410.111.49.6
max10.221.44.511.13.924.910.912.010.2
min9.519.04.210.13.521.99.510.79.0

Zapus hudsonius canadensis, St. Methode, Quebec.
4 mean9.619.24.310.13.622.69.810.99.0
max9.919.74.510.23.723.510.111.29.0
min9.118.53.99.93.321.79.510.78.7

Pancake Bay, Algoma District, Ontario.
11 mean9.618.84.110.03.522.29.610.49.2
max10.019.44.410.33.622.89.710.69.7
min9.218.33.89.63.321.89.29.88.6

Franz, Ontario.
5 mean9.819.44.210.33.522.69.810.79.0
max10.019.84.310.53.623.210.211.19.0
min9.618.94.110.23.422.19.610.48.8

Zapus hudsonius hudsonius, Fort Severn, Ontario.
4 mean9.919.34.29.93.522.79.610.99.0
max10.119.74.310.13.623.39.811.09.0
min9.819.04.09.73.422.09.310.79.0

Oxford House, Manitoba.
6 mean9.619.14.49.93.522.39.710.49.0
max9.919.84.610.13.723.110.010.89.0
min9.318.74.29.63.321.79.59.88.8

Zapus hudsonius hudsonius, Emma Lake, Saskatchewan.
2 mean9.819.44.39.93.622.79.810.99.0
max9.819.54.39.93.622.99.910.99.0
min9.819.34.29.93.522.49.610.98.9

Lac la Nonne, Alberta.
4 mean9.819.14.210.03.422.49.710.59.0
max9.819.44.210.03.522.69.910.59.0
min9.718.84.19.93.322.29.610.58.9

1 mi. NW Junct. Irons Cr. and Laird River, British Columbia.
3 mean9.619.04.39.93.522.29.710.69.0
max9.619.34.310.13.522.69.810.99.0
min9.518.74.39.73.421.79.510.38.9

Zapus hudsonius intermedius, Blackner, North Dakota.
2 mean10.119.64.310.03.622.79.911.19.0
max10.120.04.410.43.723.29.911.49.0
min10.019.14.19.83.422.29.810.89.0

Cannon Ball, North Dakota.
2 mean9.819.14.410.13.622.09.511.09.0
max9.919.14.410.23.722.49.811.39.0
min9.619.04.39.93.421.89.210.79.0

Elk River, Minnesota.
8 mean9.519.24.29.93.422.59.610.69.0
max9.719.64.310.03.623.09.911.09.0
min9.218.94.09.93.322.19.410.29.0

E Okeboji Lake, Iowa.
3 mean9.619.34.29.93.422.29.310.29.0
max9.819.34.210.03.422.29.310.29.0
min9.419.34.29.83.422.29.210.29.0

Turkey Run State Park, Indiana.
2 mean9.518.94.29.63.522.39.810.49.0
max9.618.94.49.63.622.49.810.89.0
min9.418.94.09.63.322.29.810.09.0

Zapus hudsonius intermedius, Jordan Cr., 3 mi. NE Fairmont, Ill.
5 mean9.619.44.110.13.622.610.010.79.2
max9.919.84.210.33.823.410.511.69.6
min9.318.94.010.03.421.99.510.38.9

Rib Hill, Wisconsin.
5 mean9.619.04.310.13.422.69.710.89.0
max10.019.84.410.53.623.710.211.29.3
min9.418.44.19.73.221.99.410.38.7

Lake St. Germain, Wisconsin.
5 mean9.718.94.110.13.522.59.610.59.0
max9.919.54.310.23.623.210.010.89.3
min9.518.43.99.83.321.89.010.28.5

Zapus hudsonius ladas, Northwest River, Labrador.
6 mean9.519.04.210.23.622.49.710.99.0
max9.620.04.410.33.623.210.211.09.4
min9.318.44.010.03.521.59.410.68.8

Moisie Bay, Labrador.
4 mean9.819.14.310.03.422.89.611.09.1
max9.819.74.410.03.523.510.111.39.8
min9.718.64.19.93.322.19.310.88.9

Zapus hudsonius pallidus, Mohawk Park, Oklahoma.
2 mean9.718.44.39.93.721.59.311.08.7
max10.118.84.39.93.721.89.411.08.7
min9.318.04.39.83.721.19.211.08.6

vicinity of Lawrence, Kansas.
10 mean9.218.84.49.83.421.69.710.99.0
max9.519.44.810.23.622.410.111.69.4
min8.918.14.09.33.321.09.010.38.8

2 mi. S Schuyler, Nebraska.
1 9.218.54.49.53.321.59.610.49.1

Zapus hudsonius pallidus, Valentine National Wildlife Refuge, Nebraska.
1 10.019.64.510.53.522.910.011.69.3

Zapus hudsonius preblei, Loveland, Colorado.
2 mean9.618.54.110.03.622.39.610.49.1
max9.619.04.210.03.622.49.810.69.1
min9.518.03.910.03.622.29.410.29.1

Spring Hill, 12 mi. N Laramie Peak, 6300 ft., Wyoming.
1 9.819.24.110.23.622.810.210.79.3

Zapus hudsonius tenellus, Hazelton, 959 ft., British Columbia.
2 mean9.619.44.410.13.522.99.610.99.2
max9.619.54.410.23.523.09.610.99.2
min9.619.34.310.03.422.79.610.89.2

Cottonwood P. O., British Columbia.
2 mean9.519.54.410.13.522.89.710.89.2
max9.619.64.510.23.522.89.810.99.3
min9.319.44.210.03.422.79.610.79.1

S end Swan Lake, British Columbia.
2 mean9.419.74.110.03.622.910.110.99.5
max9.419.74.210.23.623.210.210.99.6
min9.319.64.09.83.522.610.010.89.4

Indianpoint Lake, 15 mi. NE Barkerville, British Columbia.
4 mean9.618.94.19.93.421.99.510.89.0
max9.619.64.210.03.623.09.811.39.4
min9.418.34.09.83.321.39.110.28.8

LITERATURE CITED

Allen, J. A.

1894. Cranial variation in Neotoma micropus due to growth and individual differentiation. Bull. Amer. Mus. Nat. Hist., 6:233-246, 1 pl., August 3, 1894.