Omphalos: An Attempt to Untie the Geological Knot

by Philip Henry Gosse

Now each of these four hundred plates, some twelve feet in length, has grown from a minute sort of bud, in the upper jaw. Its base is hollow, resting on the formative pulp which is developed from the gum. The pulp is understood to be the immediate origin of the hairy fringe, while a dense vascular substance, seated between the bases of the plates, forms the plate itself. When the plate reaches a certain length, its diameter has become greatly attenuated, and its tip is constantly breaking away, leaving the hair projecting. There is therefore a continual disappearance of the substance of the plates at the tips, and a continual growth at the base to supply the deficiency; and even more, at least during the period of adolescence, because the actual dimensions of the plates have to be increased in the ratio of the growth of the whole animal.

Here, again, we read a record of past history. The Whale is known to be a long-lived animal; and a period of many years must have passed in bringing these plates of baleen to their present maturity. Yet the vast organism before us has been created in its vastness but to-day.

On the most prominent shelf of yonder precipice, a sharp buttress of naked limestone, stands an Ibex, guarding, like a watchful sentinel, the herd in the sheltered valley which own his leadership. The pair of noble horns, which are at once his defence and his pride, are marked throughout their ample curve with semi-rings, or knobs, on their anterior side. These afford us an infallible criterion of the animal's age.

We can count in this Ibex fourteen of such prominent bosses. Now the horn in these animals is not shed during life, but consists of a persistent sheath of horny substance, enveloping a bony core. Until full adult age, both the core of bone and the sheath of horn are continually growing; and in the spring, when there is an unusual augmentation of vital energy in the system, the increase is more than usually rapid. At this season, the new matter deposited in the corneous sheath accumulates in the form of one of these bosses, each of which is therefore produced at the interval of a year. As the first boss appears in the second year of the animal's age, we have but to add one to the number of the bosses on each horn, and we have the number of years which it has lived. The Ibex before us is just fifteen years old.

HORNS OF STAG;
In their successive developments.

Yon Stag that is rubbing his branchy honours against a tree in the glade,—can we apply the same criterion to him? Not exactly: for the horns of all the Deer-tribe are of a different structure from those of the Capradæ. They are bones of great solidity, not invested with any corneous sheath, but clothed for a certain portion of their duration with a living vascular skin, and are shed every year during life and as constantly renewed.

Yet the bony horns of the Stag are no less sure a criterion of age, at least up to a certain period—than are those of the hollow-horned Ibex. In the spring of the second year of the Fawn, the horns first appear, seated on bony footstalks that spring from the frontal bone. The skin that covers these knobs begins to swell and to become turgid with blood supplied by enlarging arteries. Layers of bone are now deposited, particle by particle, on the footstalks, with surprising rapidity, producing the budding horns, which grow day by day, still covered by the skin, which grows also in a corresponding ratio. This goes on till a simple rod of bone is formed, without any branches. When this is complete, the course of the arteries that supplied the skin is cut off by fresh osseous particles deposited in a thick ring around the base. The enveloping skin then dies, and is soon rubbed off.

After a few months, the connexion of the now dead bone with the living is dissolved by absorption, and the horns fall off.