At the time when these observations were made both the Gnus, according to Sir Andrew Smith, inhabited, in enormous multitudes, the grassy plains to the northward of the Vaal River, and after the fall of the summer rains were in the habit of advancing simultaneously in large herds as far as the southern branches of the Orange River. But on reaching the latter stream singularly enough the Brindled Gnu ceased to advance, and the Common Gnu alone passed into the Cape Colony. The appearance of the latter to the south of the Orange River was the signal for hunters of all denominations to prepare for the chase, and though the yearly slaughter was very great the herds in those days never ceased to renew their periodical visitations. Strongly expressed as are the views of both Andrew Smith and Harris as to the line of demarcation between the two species of Gnu, there seems to be some doubt as to their correctness. An excellent recent authority, Mr. H. A. Bryden (‘Kloof and Karroo’), tells us that the Brindled Gnu was in former days certainly a “denizen, albeit a rare one, south of the Orange River,” and gives us Gordon-Cumming as an authority. Gordon-Cumming asserts (‘Hunter’s Life in South Africa,’ p. 148) that he met with the Brindled Gnu in the Karroo country west of Colesberg, in what is now the Hopetown division of the Cape Colony. But Mr. Bryden admits that it has now for many years been extinct in that district. In the Transvaal, also, he tells us, the Boers have, of late years, played sad havoc with this singular Antelope, not long ago found in countless thousands on the plains of that Republic.
Messrs. Nicolls and Eglington, in their ‘Sportsman in South Africa,’ tell us the same story. Writing in 1892, they state that, except in some of the northern districts along the Crocodile River, the Brindled Gnu is now extremely scarce in the Transvaal, and practically extinct in the Orange Free State. But it is still met with in Bechuanaland, and is fairly plentiful along the edges of the Great Kalahari Desert. “In the Lake Ngami district, on both banks of the Botletle River, and from thence right up to the Chobe and Zambesi, it is quite common in suitable localities, and, at the present time, large troops may be seen on the Ma-Chara-Chara and Mababe Flats, and in the country surrounding the great salt-pans of Makari-Kari, through which the main road passes to the Victoria Falls.”
So much for the occurrence of this species in the South and South-west of Africa, past and present. But we must now trace its distribution to the north; for it is an extraordinary fact that the Brindled Gnu, instead of being confined, like its sister species, to a small part of South Africa, extends up along the eastern coast certainly as far as Kilimanjaro, and perhaps even into Sennaar, where reports of the occurrence of a Gnu-like Antelope were made to Heuglin. It is, however, possible that some of these northern Gnus may belong to the White-maned species, which we shall presently mention.
Beginning from the Limpopo, Mr. Selous tells us that the Brindled Gnu is found all over this portion of South-eastern Africa up to the Zambesi, in districts suitable to its habits—that is, “in open downs devoid of bush and in open glades in the forest,” but not in hilly countries. Peters and Sir John Kirk both enroll it among the mammals of Zambesia, the latter author stating that at the time of his visit it was “very abundant in considerable herds in the Batoka country, also near Lake Shirwa, and at Shupanga on the Zambesi.” As to its present existence in the Shiré Highlands, Mr. B. L. Sclater, R.E., sends us the following notes.—“In November 1891, while travelling between Zomba and Milanji, I was shooting in the marshes on the west bank of the Tochila. I saw a Gnu and tried to stalk it, but it was right in the open and I could not get near it. My head man, a Swahili, said it was a Gnu (Nyumbo), and he was well acquainted with that animal. Again, in 1892, at Midima, to the south of the Tochila, I obtained a tail of a Gnu from a native, who told me that formerly there was a large herd of them on these plains, but that he thought they were now all killed. I believe that Mr. Sharpe has met with the Gnu on the plains to the west of the Upper Shiré, between Matope and the sources of the Lesungwi River.”
As regards the more northern portion of the British Central African Protectorate, Mr. Crawshay tells us that the Gnu is apparently unknown to the natives round the northern half of Lake Nyasa, and is not met with anywhere in the immediate neighbourhood of the Lake, though found a little to the south-east, and also, he believes, to the south-west.
In 1864 Speke met with the Brindled Gnu in large herds in Khutu, on the western borders of Uzaramo, close to the Kingani River, where it inhabits the “park-like lands adjoining the stream.”
Later on Sir John Kirk obtained heads of it in the same district, and has favoured us with the following notes on its occurrence in this part of German East Africa:—
“As regards the Brindled Gnu in East Africa, I may say that, although familiar with this animal on the Upper Zambezi near the Victoria Falls, where they were common in 1860, I have only since shot them in Ukami, to the west of Dar-es-salam, and on the River Wami inland from Bagomoyo. In the plains and on the rolling ground between the River Rufiji and the River Wami they used to be common. I have shot them within ten miles of the coast, and I believe that they extend back to the foot of the mountains.
“As I was not then acquainted with the species or variety lately found by Jackson, I cannot from memory express any opinion as to the identity of these animals, further than that they seemed to me identical with the Brindled Gnus I had killed years previously near the Victoria Falls and Sesheke on the Upper Zambezi.
“In East Africa, near the coast, in the places above-named where I found this Antelope, it never occurs in numbers, but is often associated as an attendant on other game, especially upon Lichtenstein’s Hartebeest. It often causes annoyance to the sportsman by giving warning of his approach to the other Antelopes.