The History of the European Fauna

by R. F. Scharff

The sequence of events in the British Islands was probably as follows:—The first comers were the members of that fauna which issued from South-western Europe; then came the Alpine, and at the same time probably the Arctic and the Oriental; and finally the Eastern or Siberian. The migrations of all but the last continued, uninterruptedly, for very long periods.

The study of these migrations has convinced me that, though climate was a powerful factor in the evolution or history of the European fauna, the geographical changes which took place on our continent in later Tertiary times exerted a yet stronger influence. The principal climatic disturbance is generally supposed to have been the so-called "Ice Age." So firmly rooted is the conviction, among naturalists of the present day, of the enormous destruction which this period produced on our European fauna, so that all animal life practically disappeared from large areas of our continent, that it is desirable that we should now shortly review the history of that remarkable period in order to ascertain in how far these views are corroborated by facts. Frequent reference, moreover, will be made throughout this work to the theories connected with the Glacial period.

It has been stated by an eminent geologist that during part of the Glacial period the climate was such that neither plants nor animals could have existed in the British Islands. If that had been so, it is evident that very few organisms could have even survived in France, though a number of Arctic species might have dragged on an existence in Southern Europe. At any rate, on the return of more genial conditions, the Arctic species would undoubtedly have been the first to gain admission to the British Islands, to re-people the arid wastes. Our supposition that the Lusitanian element in the British fauna is the oldest would therefore be wrong. From early Tertiary times onward, the climate of Europe, which was then semi-tropical, gradually became more and more temperate; until finally the Ice Age or Glacial period arrived, during which, according to Professor J. Geikie—one of our highest authorities on this subject—a great part of Northern Europe became practically uninhabitable owing to the severity of the climate.

To enable us to judge better of the true value of the many hypotheses which have been advanced to account for this supposed extraordinary fall of temperature during the "Ice Age," we must compare the views of other authorities with the one just quoted. I do not propose to discuss the causes which have led to the production of the Glacial period—those interested in these questions should consult the writings of Dr. Croll, Professor J. Geikie, Professor Bonney, Mr. Falsan, and others—but merely to give the climatic aspects from a physical, zoological, and botanical point of view.

According to Professor Penck (a, p. 12), the nature of the glacial climate can be determined by comparing the snow-line of the Glacial period with that of the present day. The position of the snow-line is dependent on two climatic factors—viz., precipitation and temperature. We know the height at which snow must have lain permanently during the Glacial period, or during the maximum phase of glaciation. If the Ice Age had been produced solely by an increase of snowfall, as has been suggested, Professor Penck tells us that then it must have snowed three or four times as much as it does now. But he does not adopt the view that the Ice Age is due to an increase of snowfall alone. His calculations, based upon the height of the snow-line, tend to show that a general decrease of temperature to the extent of from 4-5 degrees Centigrade (all other atmospheric conditions remaining the same as now) would be sufficient to give us back the Glacial period.

Professor Neumayr (p. 619) adopted a similar principle in determining the temperature which prevailed in Europe during the Glacial period. Snow now lies in the Pyrenees 1000 metres higher than it did then, 1,200 metres higher in the Alps, and 800 metres higher in the Tatra mountains. Since the temperature in Central Europe decreases by half a degree Centigrade for every 100 metres of elevation, it follows that if the glacial phenomena had only been brought about by a decrease of temperature without an increase of moisture, we should have had a reduction of temperature during the Glacial period of six degrees Centigrade in the Pyrenees, of seven degrees in the Alps, and of four in the Tatra mountains. The general lowering of the temperature of Europe, says Professor Neumayr, could not have amounted to more than six degrees Centigrade. Moreover, he is of opinion that the very low snow-line in the British Islands proves that even during the Ice Age a comparatively mild climate prevailed there, and that the climatic conditions generally, in the different parts of Europe, were relatively about the same as they are now.

Professor J. Geikie does not give us his views as to the temperature of the Glacial period, but he maintains that a lowering of the temperature is evinced not only by the widespread phenomena of glaciation, but by the former presence in our temperate latitudes of a northern fauna and flora.

Mr. Charles Martins, who based his calculations on the temperature during the Glacial period on the glaciers of Chamounix, concluded that it only needed a lowering of the temperature to the extent of four degrees Centigrade to bring the glaciers down to the plain of Geneva, and in fact give us back the Glacial period. It need not surprise us, therefore, that the French geologist, Mr. Falsan, the author of La période glacière, is of opinion (p. 230) that the mean annual temperature of France during the Glacial period was approximately from 6-9 degrees Centigrade, perhaps more. Close to the immense glaciers of the Rhone, it might have been about six degrees. This is the actual mean annual temperature of the South-west of Sweden and Norway, or the North of Scotland.

Although all these investigations tend to show that the climate of Europe during the Glacial period was by no means so severe as we are often led to believe, yet there exists also a school of geologists who maintain there was actually a higher temperature than at present. The inconsistency of mentioning heat in connection with ice and snow is more apparent, however, than real, for we must remember Tyndall's original remark on this subject. It is the snow, he says, which feeds the glaciers. But the snow comes from the clouds, and these again originate from the vapours which the sun causes to be absorbed from the ocean. Without the sun's heat, we should have no water vapour in the atmosphere; without vapour, no clouds; without clouds, no snow; without snow, no glaciers. The ice of glaciers, therefore, owes its origin indirectly to the sun's heat. It has been supposed that if the sun's heat diminished, larger glaciers would form than those existing to-day, but the diminution of the solar heat would infallibly reduce the amount of water vapour in the air, and would thus stop the very source of glaciers.

Mr. Falsan even admits that without a change of the mean annual temperature (p. 201) of Europe, the central portions of our continent might at this period have enjoyed an insular climate. This more equable and humid climate could, within certain limits, favour the development of the ancient glaciers by increasing the snowfall and slackening the summer rate of melting.