Origin: sagittal crest; anteriorly with internal temporalis from frontal bone; posteriorly with internal temporalis from interparietal bone.

Insertion: lateral surface of coronoid process of mandible.

3. Internal temporalis

Origin: sagittal crest and skull roof, including posterior two-thirds of frontal bone, whole of parietal, and dorsalmost portions of squamosal and alisphenoid.

Insertion: medial surface of coronoid process; dorsal edge of coronoid process.

Fig. 8. Didelphis marsupialis. Showing masseter and temporal muscles. Skull KU 3780, 1 mi. N Lawrence, Douglas Co., Kansas. × 3/5.

Temporal Openings

In discussions of the morphology and functions of the adductor mechanism of the lower jaw, the problem of accounting for the appearance of temporal openings in the skull is often encountered. Two patterns of explanation have evolved. The first has been the attempt to ascribe to the constant action of the same selective force the openings from their inception in primitive members of a phyletic line to their fullest expression in terminal members. According to this theory, for example, the synapsid opening appeared originally to allow freer expansion of the adductor muscles of the jaw during contraction, and continued selection for that character caused the openings to expand until the ultimately derived therapsid or mammalian condition was achieved.