Natural History of the Mammalia of India and Ceylon

by Robert Armitage Sterndale

HABITAT.—The hilly parts Moupin, Eastern Thibet.

Ailuropus melanoleucos.

DESCRIPTION.—The Ailuropus has a thick-set heavy form. His head is short, rather slender in front, but extremely enlarged in the middle and after part; the nose is small and naked at its extremity; the forehead very large and convex; the eyes are small; the ears short, wide between and rounded at the ends; neck thick and very strong; the body is squat and massive; the tail is so short as to be hardly distinguishable. The feet are short, very large, nearly of the same length, terminated by five toes very large and with rounded ends, the general conformation of which recalls in all respects those of the bears, but of which the lower parts, instead of being completely placed on the sole in walking and entirely naked or devoid of hair, are always in great measure raised, and abundantly clad with fur to almost their full extent.

On the hind feet can be noticed at the base of the toes a transverse range of five little fleshy pads, and towards the anterior extremity of the metatarsal region another naked cushion placed transversely; but between these parts, as well as the posterior two-thirds of the planta, the hair is as abundant and as long almost as on the upper part of the foot. In the fore-limbs the disposition is much the same, though the metacarpal cushion may be larger; and there is another fleshy pad without hair near the claws.

The Ailuropus is thus an animal not strictly plantigrade, like the Bears in general, or the same as the Polar Bear, of which the feet, although placed flat on the earth, are not devoid of hair; but, on the contrary, the Ailuropus resembles the Ailurus, which is semi-plantigrade, yet hairy under Its soles.

The colouring of the Ailuropus is remarkable: it is white with the exception of the circumferences of the eyes, the ears, the shoulders, and the lower part of the neck which are entirely black. These stand out clearly on a groundwork of slightly yellowish-white; the spots round the eyes are circular, and give a strange aspect to the animal; those on the shoulders represent a sort of band placed transversely across the withers, widening as they descend downwards to lower limbs. The hinder limbs are also black from the lower part of the thigh down to the toes, but the haunches, as also the greater part of the tail, are as white as the back and belly; the colouring is the same in young and old. The fur is long, thick, and coarse, like that of the bears.

From the general form of the skull it would seem impossible to determine the family to which this animal belongs. In effect the head differs considerably from the Ursidæ and the Mustelidæ, and presents certain resemblances to that of the hyæna; but there are numerous and important particulars which indicate a special zoological type, and it is only by an inspection of the dental system that the natural affinities of the Ailuropus can be determined.

In the upper jaw the incisors are, as usual, in three pairs. They are remarkable for their oblique direction; the centre ones are small and a little widened at the base; the second pair are stronger and dilated towards the cutting edge; the external incisors are also strong and excavated outside to admit the canines of the lower jaw. The canines are stout, but short, with a well-marked blunt ridge down the posterior side, as in the Malayan bears.

The molars are six in number on each side, of which four are premolars, and two true molars. The first premolar, situated behind, a little within the line of the canine, is very small, tuberculiform, and a little compressed laterally. The second is strong and essentially carnassial; it is compressed laterally and obliquely placed. It is furnished with three lobes: the first lobe is short, thick, and obtuse; the second is raised, triangular and with cutting edges; the third of the size of the first, but more compressed—in short, a double-fanged tooth. This molar differs considerably from the corresponding tooth of the bear by its form and relative development, since in that family it is one-fanged, very low and obtuse. On the contrary, it approaches to that of the hyænas and felines. With the panda (Ailurus fulgens) the corresponding premolar is equally large, double-fanged and trenchant, but the division in lobes is not so marked.