The same law of development presides over the vegetable kingdom. Amongst phanerogamous plants, a certain number of organs appear to be always present, either in a developed or rudimentary state; and those which are rudimentary can be developed by cultivation. The flowers which bear stamens on one stalk and pistils on another, can be caused to produce both, or to become perfect flowers, by having a sufficiency of nourishment supplied to them. So also, where a special function is required for particular circumstances, nature has provided for it, not by a new organ, but by a modification of a common one, which she has effected in development. Thus, for instance, some plants destined to live in arid situations, require to have a store of water which they may slowly absorb. The need is arranged for by a cup-like expansion round the stalk, in which water remains after a shower. Now the pitcher, as this is called, is not a new organ, but simply a metamorphose of a leaf.
These facts clearly shew how all the various organic forms of our world are bound up in one—how a fundamental unity pervades and embraces them all, collecting them, from the humblest lichen up to the highest mammifer, in one system, the whole creation of which must have depended upon one law or decree of the Almighty, though it did not all come forth at one time. After what we have seen, the idea of a separate exertion for each must appear totally inadmissible. The single fact of abortive or rudimentary organs condemns it; for these, on such a supposition, could be regarded in no other light than as blemishes or blunders—the thing of all others most irreconcilable with that idea of Almighty Perfection which a general view of nature so irresistibly conveys. On the other hand, when the organic creation is admitted to have been effected by a general law, we see nothing in these abortive parts but harmless peculiarities of development, and interesting evidences of the manner in which the Divine Author has been pleased to work.
We have yet to advert to the most interesting class of facts connected with the laws of organic development. It is only in recent times that physiologists have observed that each animal passes, in the course of its germinal history, through a series of changes resembling the permanent forms of the various orders of animals inferior to it in the scale. Thus, for instance, an insect, standing at the head of the articulated animals, is, in the larva state, a true annelid, or worm, the annelida being the lowest in the same class. The embryo of a crab resembles the perfect animal of the inferior order myriapoda, and passes through all the forms of transition which characterize the intermediate tribes of crustacea. The frog, for some time after its birth, is a fish with external gills, and other organs fitting it for an aquatic life, all of which are changed as it advances to maturity, and becomes a land animal. The mammifer only passes through still more stages, according to its higher place in the scale. Nor is man himself exempt from this law. His first form is that which is permanent in the animalcule. His organization gradually passes through conditions generally resembling a fish, a reptile, a bird, and the lower mammalia, before it attains its specific maturity. At one of the last stages of his fœtal career, he exhibits an intermaxillary bone, which is characteristic of the perfect ape; this is suppressed, and he may then be said to take leave of the simial type, and become a true human creature. Even, as we shall see, the varieties of his race are represented in the progressive development of an individual of the highest, before we see the adult Caucasian, the highest point yet attained in the animal scale.
To come to particular points of the organization. The brain of man, which exceeds that of all other animals in complexity of organization and fulness of development, is, at one early period, only “a simple fold of nervous matter, with difficulty distinguishable into three parts, while a little tail-like prolongation towards the hinder parts, and which had been the first to appear, is the only representation of a spinal marrow. Now, in this state it perfectly resembles the brain of an adult fish, thus assuming in transitu the form that in the fish is permanent. In a short time, however, the structure is become more complex, the parts more distinct, the spinal marrow better marked; it is now the brain of a reptile. The change continues; by a singular motion, certain parts (corpora quadragemina) which had hitherto appeared on the upper surface, now pass towards the lower; the former is their permanent situation in fishes and reptiles, the latter in birds and mammalia. This is another advance in the scale, but more remains yet to be done. The complication of the organ increases; cavities termed ventricles are formed, which do not exist in fishes, reptiles, or birds; curiously organized parts, such as the corpora striata, are added; it is now the brain of the mammalia. Its last and final change alone seems wanting, that which shall render it the brain of MAN.” [201] And this change in time takes place.
So also with the heart. This organ, in the mammalia, consists of four cavities, but in the reptiles of only three, and in fishes of two only, while in the articulated animals it is merely a prolonged tube. Now in the mammal fœtus, at a certain early stage, the organ has the form of a prolonged tube; and a human being may be said to have then the heart of an insect. Subsequently it is shortened and widened, and becomes divided by a contraction into two parts, a ventricle and an auricle; it is now the heart of a fish. A subdivision of the auricle afterwards makes a triple-chambered form, as in the heart of the reptile tribes; lastly, the ventricle being also subdivided, it becomes a full mammal heart.
Another illustration here presents itself with the force of the most powerful and interesting analogy. Some of the earliest fishes of our globe, those of the Old Red Sandstone, present, as we have seen, certain peculiarities, as the one-sided tail and an inferior position of the mouth. No fishes of the present day, in a mature state, are so characterized; but some, at a certain stage of their existence, have such peculiarities. It occurred to a geologist to inquire if the fish which existed before the Old Red Sandstone had any peculiarities assimilating them to the fœtal condition of existing fish, and particularly if they were small. The first which occur before the time of the Old Red Sandstone, are those described by Mr. Murchison, as belonging to the Upper Ludlow Rocks; they are all rather small. Still older are those detected by Mr. Philips, in the Aymestry Limestone, being the most ancient of the class which have as yet been discovered; these are so extremely minute as only to be distinguishable by the microscope. Here we apparently have very clear demonstrations of a parity, or rather identity, of laws presiding over the development of the animated tribes on the face of the earth, and that of the individual in embryo.
The tendency of all these illustrations is to make us look to development as the principle which has been immediately concerned in the peopling of this globe, a process extending over a vast space of time, but which is nevertheless connected in character with the briefer process by which an individual being is evoked from a simple germ. What mystery is there here—and how shall I proceed to enunciate the conception which I have ventured to form of what may prove to be its proper solution! It is an idea by no means calculated to impress by its greatness, or to puzzle by its profoundness. It is an idea more marked by simplicity than perhaps any other of those which have explained the great secrets of nature. But in this lies, perhaps, one of its strongest claims to the faith of mankind.
The whole train of animated beings, from the simplest and oldest up to the highest and most recent, are, then, to be regarded as a series of advances of the principle of development, which have depended upon external physical circumstances, to which the resulting animals are appropriate. I contemplate the whole phenomena as having been in the first place arranged in the counsels of Divine Wisdom, to take place, not only upon this sphere, but upon all the others in space, under necessary modifications, and as being carried on, from first to last, here and elsewhere, under immediate favour of the creative will or energy. [204] The nucleated vesicle, the fundamental form of all organization, we must regard as the meeting-point between the inorganic and the organic—the end of the mineral and beginning of the vegetable and animal kingdoms, which thence start in different directions, but in perfect parallelism and analogy. We have already seen that this nucleated vesicle is itself a type of mature and independent being in the infusory animalcules, as well as the starting point of the fœtal progress of every higher individual in creation, both animal and vegetable. We have seen that it is a form of being which electric agency will produce—though not perhaps usher into full life—in albumen, one of those compound elements of animal bodies, of which another (urea) has been made by artificial means. Remembering these things, we are drawn on to the supposition, that the first step in the creation of life upon this planet was a chemico-electric operation, by which simple germinal vesicles were produced. This is so much, but what were the next steps? Let a common vegetable infusion help us to an answer. There, as we have seen, simple forms are produced at first, but afterwards they become more complicated, until at length the life-producing powers of the infusion are exhausted. Are we to presume that, in this case, the simple engender the complicated? Undoubtedly, this would not be more wonderful as a natural process than one which we never think of wondering at, because familiar to us—namely, that in the gestation of the mammals, the animalcule-like ovum of a few days is the parent, in a sense, of the chick-like form of a few weeks, and that in all the subsequent stages—fish, reptile, &c.—the one may, with scarcely a metaphor, be said to be the progenitor of the other. I suggest, then, as an hypothesis already countenanced by much that is ascertained, and likely to be further sanctioned by much that remains to be known, that the first step was an advance under favour of peculiar conditions, from the simplest forms of being, to the next more complicated, and this through the medium of the ordinary process of generation.
Unquestionably, what we ordinarily see of nature is calculated to impress a conviction that each species invariably produces its like. But I would here call attention to a remarkable illustration of natural law which has been brought forward by Mr. Babbage, in his Ninth Bridgewater Treatise. The reader is requested to suppose himself seated before the calculating machine, and observing it. It is moved by a weight, and there is a wheel which revolves through a small angle round its axis, at short intervals, presenting to his eye successively, a series of numbers engraved on its divided circumference.
Let the figures thus seen be the series, 1, 2, 3, 4, 5, &c., of natural numbers, each of which exceeds its immediate antecedent by unity.