Range.—In Canada, southern Ontario and Quebec; in the United States, northwestern Vermont and western New York south to northern Florida, east to central Montana, eastern Wyoming and Colorado, and New Mexico; introduced into the Colorado River system of California, Nevada, Arizona and New Mexico; in México, the northern part of the states of Tamaulipas, Nuevo León, Coahuila, and eastern Chihuahua (see map, [Fig. 19]).

Diagnosis.—Juvenal pattern uniform tan or brownish lacking markings, having whitish dots or spots, or having well-defined, blackish ocelli or spots; surface of carapace "sandpapery" in adult males; conical projections (in some subspecies) along anterior edge of carapace in large females; contrasting pattern of blackish marks on pale background (in some subspecies) on dorsal surface of limbs of adult males.

Opisthotic-exoccipital spur well-developed; epiplastral callosity, when present, not covering entire surface.

Description.—Septal ridges present; external and proportional characteristics variable (see accounts of subspecies); range in length of plastron (cm.) of ten [487] largest specimens of each sex (mean follows extremes), males, 13.8-16.0, 14.4; females, 26.0-31.0, 28.0.

Fig. 19. Geographic distribution of Trionyx spinifer.

Guide to subspecies:

  1. T. s. spinifer
  2. T. s. hartwegi
  3. T. s. asper
  4. T. s. pallidus
  5. T. s. guadalupensis
  6. T. s. emoryi

Greatest width of skull usually at level of squamosal (74%); foramen magnum rhomboidal; ventral surface of supraoccipital spine narrow proximally, usually having medial ridge; opisthotic-exoccipital spur well-developed (66%); distal part of opisthotic wing tapered, not visible in dorsal view; lateral condyle of articular surface of quadrate larger than medial articular surface, not tapered posteriorly; maxillaries in contact above premaxillaries (88%); usually a combination of seven neurals, seven pairs of pleurals and contact of seventh pair of pleurals (83%); angle of epiplastron approximately 90 degrees; callosities when present on epiplastron not covering entire surface; hyo-hypoplastral suture usually present.