Trionyx muticus muticus Webb, Publ. Mus. Nat. Hist. Univ. Kansas, 11:520, August 14, 1959.

Potamochelys? microcephala Gray, Proc. Zool. Soc. London, p. 87, 1864.

Type.—Lectotype, Museum d'Histoire Naturelle, Paris, No. 8813; dried carapace and plastron; obtained from the Wabash River, New Harmony, Posey County, Indiana, by C. A. Lesueur in August, 1827 ([Pl. 53]).

Range.—Central United States; in the Mississippi River drainage from extreme western Pennsylvania, southern Minnesota and South Dakota south to Tennessee, Louisiana and Oklahoma; streams of the Gulf Coast drainage from the Mississippi River in Louisiana westward into Texas including the Colorado River drainage (see map, [Fig. 22]).

Diagnosis.—Juvenal pattern of dusky dots and usually short lines or bacilliform marks; ill-defined pale stripes on snout usually evident just in front of eyes; pale postocular stripe lacking thick, black borders that are approximately one-half width of pale stripe (except some in the Colorado River drainage of Texas).

Description.—Plastral length of smallest hatchling, 2.1 centimeters (INHS 3458); of largest male, 14.0 centimeters (CNHM 92003); of largest female, 21.5 centimeters (KU 2308).

Juvenal pattern of dusky, grayish marks lacking sharp margins, and usually consisting of both small spots and short streaks or dashes, the former predominating; short streaks or dashes occasionally lacking (TU 14375, [Pl. 45], bottom, left; UMMZ 92751); markings variable in number, few and widely spaced, or several and closely approximated ([Pl. 45], top, topotypes); pale rim [535] separated from ground color by ill-defined, dusky margin; pattern on adult males well-defined resembling that of hatchlings (TU 16172.1, 16173), scarcely discernable (TU 13294), or absent (TU 1242); mottled and blotched pattern on carapace usually contrasting in large females.

Pale stripes extending forward from eyes usually not more than half distance to tip of snout; inner borders of pale stripes on snout usually absent or dusky and indistinct, occasionally blackish (TU 14606); outer borders of pale stripes darker than inner borders, usually blackish; pale stripes on snout occasionally absent (CNHM 7845, UMMZ 92665, TU 5989, none of these specimens being large females); pale postocular stripe having narrow, dusky or blackish borders (especially UMMZ 92751, TU 14436); pale postocular stripe usually complete, occasionally interrupted having prominent dark-bordered anterior segment just behind eye (TU 14416); lower border of postocular stripe usually in contact with dusky postlabial line; no other markings on side of head; pattern on dorsal surface of soft parts of body not contrasting, composed of closely approximated fine markings that are little darker than background, over-all coloration pale grayish; occasionally, few larger and more contrasting markings on hind limbs (UMMZ 92751, TU 14436).

Underparts white, usually lacking markings; occasional dusky markings on plastral area (UMMZ 110502), dark spots or flecks on undersurface of carapace (BCB 6043, UMMZ 92666), or markings on throat (UMMZ 95032).

Surface of carapace smooth in adult males; large females lacking prominences posteriorly in center of carapace or in nuchal region; anterior edge of carapace smooth in both sexes, but occasionally having regularly spaced furrows or wrinkles ([Fig. 8g]).