The skull is in the form of a narrow oval, sharply narrowed posteriorly and truncate. The orbits are located well forward and their posterior border lies in front of the line dividing the skull transversely into equal parts. The interorbital space is greater than the diameter of the orbit. Impressions of teeth are preserved on the premaxillæ and maxillæ; there are 8 of them in a distance of 3 mm. The teeth appear to be mere blunt denticles and were possibly pleurodont.
The elements of the cranium are very poorly preserved. It has been impossible to determine all of the sutures. The bones of the premaxillary region have been destroyed, but the arrangement of them was probably not far different from that which obtains in other members of the genus. The posterior boundaries of the nasals are preserved and prove this element to have had an obtuse posterior border. The sutures bounding the frontals are clear and show that they were small and that they formed a part of the inner boundary of the orbits. The parietal is recognized as a large element, apparently the largest in the skull. Together the parietals form a wide oval inclosing, on the median suture, the circular pineal foramen. The parietals are sculptured with coarse radiating grooves and ridges, much after the manner of Erpetosaurus radiatus Cope. The pittings present on that form are, however, absent in T. minimus. The sutures bounding the postparietal are tolerably well defined and these show that element to have been rather large and quadrate, with the usual relations. The tabulare is distinct, triangular, and small. It is produced into an angle on the posterior border strongly recalling a similar condition in T. punctulatus. The boundaries of the prefrontals and the upper borders of the maxillæ are not clearly ascertained. The lacrimal has not been detected. The post-frontal and postorbital form the posterior boundary of the orbit, although all of the limits of the latter element have not been definitely determined. The position of the supratemporal is well assured, although its entire boundaries are not determined. It has the usual relations and joins the parietal broadly. The jugal is broad and widens posteriorly to join the squamosal, which, as usual, forms the quadrate angle of the skull. The sutures bounding the quadratojugal and the posterior end of the maxilla are not determined.
There are but two fragmentary vertebræ preserved and an estimate based on the length of these remains gives about 30 presacral vertebræ. The structure of the vertebræ preserved can not be ascertained, but the neural spines appear to have been low and stout.
There are six elements of the pectoral girdle preserved. These are: the six clavicles, the interclavicle, the coracoid of one side, and the two scapulæ. The interclavicle is rhomboid in form and acuminate posteriorly. It is sculptured with radiating grooves and ridges. It is quite different from the same element in T. punctulatus, in that the base is acuminate, not truncate. The clavicle presents much the same shape as does that element in Erpetosaurus tabulatus. It is ornamented by a sculpturing of radiating lines which take their origin from the lower external angle as the bone lies in the matrix. The clavicle is somewhat triangular in shape and lies close to the skull, but this close approximation of the pectoral elements to the cranium is due probably to post-mortem shifting, since the scapulæ are shifted backward. There can be little doubt, however, that the pectoral arch was close to the cranium. There is an oval fragment preserved on the left of the specimen which I take to be a portion of the coracoid. The scapula is preserved entire on the left side and is represented by fragments on the right side. It is almost semicircular in form and narrows externally until it is somewhat fan-shaped. There appears to be an ornamentation of lines on the surface of the bone. These lines follow the contour of the anterior border.
The arm is preserved nearly complete on the left side, and the right side shows the humerus and the forearm. The humeri are unusual in having well-developed articular ends as though the endochondral tissue was well developed. The humerus is expanded at the ends and it is larger at the upper than at the lower end. The ulna is expanded at the proximal extremity, but is more attenuated at the distal portion. It is shorter than the humerus by about one-third of its own length. The radius is a mere slender rod of bone and presents well-developed articular ends. It is slightly shorter than the ulna. The carpus is unossified and its position is represented by a blank space. There are phalanges of 4 digits preserved and they are 4 in number. The phalangeal elements, like the other bones of the extremity, have the articular surfaces prominent, with the terminal phalanx claw-like.
There are no ribs nor traces of them preserved, and a conjecture as to their character can not be hazarded, since they are known in but two other species, in which they are slender and curved. There is no evidence of a ventral scutellation, and so far as is at present known this structure is absent from all of the species of the genus, or at least it is weakly developed.
The ilium is all that is preserved of the pelvis. The bone itself has disappeared and has left a depression which shows this element to have been an elongate rod very similar to that described for Micrerpeton. The sacral vertebra seems to be indicated by a depression between the iliac depressions.
One hind limb is preserved nearly entire and the greater part of the other is also preserved, although some of the phalanges are disturbed. The femur is slender and more elongate than the humerus. It has well-formed, rounded, articular ends. The tibia presents unusual characters in that its ends are truncate, as though the cartilage composing its articular surfaces was not so highly calcified as in the other limb bones. It is somewhat expanded at the ends and is throughout its length broader than the femur. The fibula, like the tibia, is a slender rod of bone, although it is somewhat shorter than the tibia. The tarsus is unossified and its position is occupied by a blank space. Portions of both feet are preserved, but only one digit in the right foot is complete. The metatarsals are elongate and -slightly expanded at the ends. There are 4 phalanges present in the complete digit, which possibly represents the fourth. The first digit is wanting, with the only terminal phalanx preserved claw-like.
| Measurements of the Type of Tuditanus minimus Moodie. | |
| mm. | |
| Median length of skull | 15 |
| Width of skull at posterior border | 16 |
| Length of orbit | 3.5 |
| Width of orbit | 2 |
| Interorbital width | 2.5 |
| Length of clavicle | 6 |
| Width of clavicle, maximum | 3.5 |
| Length of interclavicle, estimated | 5 |
| Width of interclavicle | 3.5 |
| Length of scapula | 3.5 |
| Width of scapula, maximum | 2.5 |
| Length of coracoid (?) | 2 |
| Length of humerus | 4 |
| Length of radius and ulna | 3 |
| Length of metacarpal | 1 |
| Length of ilium | 2.5 |
| Length of femur | 4.5 |
| Length of tibia and fibula | 3 |
| Length of foot | 3.5 |
| Length of metatarsal | .75 |
Tuditanus walcotti Moodie.