In seeking to define our position in regard to the theory of descent it is most important that we should recognise that, when it is looked into closely, the true problem at issue is not a special zoological one, but is quite general, and also that it is not a new growth which has sprung up suddenly and found us unprepared, but that it is very ancient and has long existed in our midst. In the whole theory the question of “descent” is after all a mere accessory. Even if it fell through and were seen to be scientifically undemonstrable, “evolution in the realm of life” would remain an indisputable fact, and with it there would arise precisely the same difficulties for the religious interpretation of the world which are usually attributed to the Theory of Descent.

Evolution or development has been a prominent idea in the history of thought since the time of Aristotle, but descent is, so to speak, a modern upstart. According to long-established modes of thought, to evolve means to pass from δυνάμει to ἰνεργεία εἴναι, from potentia to actus, from the existence of the rudiment as in the seed to full realisation as in the tree. In the course of its [pg 129] development the organism passes through many successive phases, which are related to one another like steps, each rising directly from the one beneath, and preparing for the one above. Thus all nature, and especially the realm of life, implies a ladder of “evolution.” What is “potentially” inherent in the lowest form of life has in the highest, as in man, become actual or “realised” through a continuous sequence of phases, successively more and more evolved. This view in its earlier forms was very far from implying that each higher step was literally “descended” from the one below it, through the physical and mental transformation of some of its representatives. As the world, in Aristotle's view for instance, had existed from all eternity, so also had the stages and forms of life, each giving rise again to its like. Indeed, the essential idea was that each higher step is simply a development, a fuller unfolding of the lower stage, and finally that man was the complete realisation of what was potentially inherent in the lowest of all.

This doctrine of evolution was in modern times the fundamental idea of Leibnitz and Kant, of Goethe, Schelling and Hegel. It brought unity and connectedness into the system of nature, united everything by steps, denied the existence of gaping chasms, and proclaimed the solidarity of all the forms of life. But to all this the idea of actual descent was unnecessary. An actual material variation and transition from one stage to another seemed to it a wooden and gross [pg 130] expression of the evolution idea, an “all too childish and nebulous hypothesis” (Hegel).

All the important results of comparative morphology and physiology, which the modern supporters of the doctrine of descent so confidently utilise as arguments in its favour, would have been welcomed by those who held the original and general evolution idea, as a corroboration of their own standpoint. And as a matter of fact they all afford conclusive proofs of evolution; but not one of them, including even the fundamental biogenetic law and the inoculated chimpanzee, is decisive in regard to descent. This contention is sufficiently important to claim our attention for a little. Let us take the last example. Transfusion of blood between two species is possible, not necessarily because they are descended from one another or from a common root, but solely because of their systematic (ideal) relationship, that is to say because they are sufficiently near to one another and like one another in their physiological qualities and functions. If, assuming descent, this homology were disturbed, and the systematic relationship done away with, for instance through saltatory evolution, the mere fact of descent would not bring the two species any nearer one another. Thus the case proves only systematic relationship, and only evolution. But as to the meaning of this systematic relationship, whether it can be “explained” by descent, whether it has existed from all eternity, or how it has arisen, the experiment does not inform us.

The same idea may be illustrated in regard to Weismann's “predicting.” This, too, is a proof of evolution, but not of descent. Exactly as Weismann predicted the striping of the hawk-moth caterpillars and the human os centrale, Goethe predicted the formation of the skull from modified vertebræ, and the premaxillary bone in man. In precisely the same way he “derived” the cavities in the human skull from those of the animal skull. This was quite in keeping with the manner and style of his Goddess Nature and her creative transformations, raising the type of her creations from stage to stage, developing and expanding each new type from an earlier one, yet keeping the later analogous to and recapitulative of the earlier, recording the earlier by means of vestigial and gradually dwindling parts.

But what has all this to do with descent? Even the “biogenetic law” itself, especially if it were correct, would fit admirably into the frame of the pure evolution idea. For it is quite consistent with that idea to say that the higher type in the course of its development, especially in its embryonic stages, passes through stages representative of the forms of life which are below it and precede it in the (ideal) genealogical tree. Indeed, the older doctrine of evolution took account of this long ago.

“The same step-ladder which is exhibited by the whole animal kingdom, the steps of which are the different races and classes, with at the one extreme the [pg 132] lowliest animals and at the other the highest, is exhibited also by every higher animal in its development, since from the moment of its origin until it has reached its full development it passes through—both as regards internal and external organisation—the essentials of all the forms which become permanent for a lifetime in the animals lower than itself. The more perfect the animal is, the longer is the series of forms it passes through.”

So J. Fr. Meckel wrote in 1812 in his “Handbook of Pathological Anatomy,” with no thought of descent. And the facts which led to the construction of the biogenetic law were discovered in no small measure by Agassiz, who was an opponent of the doctrine of descent.[31]

But the advance from the doctrine of evolution to that of descent was imperatively prompted by a recognition of [pg 133] the fact that the earth is not from everlasting, and that the forms of life upon it are likewise not from everlasting, that, in fact, their several grades appear in an orderly ascending series. It is therefore simpler and more plausible to suppose that each higher step has arisen from the one before it, than to suppose that each has, so to speak, begun an evolution on its own account. A series of corroborative arguments might be adduced, and there is no doubt, as we have said before, that the transition from the general idea of evolution to that of descent will be fully accomplished. But it is plain that the special idea of descent contributes nothing essentially new on the subject.

It is an oft-repeated and self-evident statement, that it is in reality a matter of entire indifference whether man arose from the dust of the earth or from living matter already formed, or, let us say, from one of the higher vertebrates. The question still would be, how much or how little of any of them does he still retain, and how far does he differ from all? Even if there be really descent, the difference may quite as well be so great—for instance, through saltatory development—that man, in spite of physical relationship, might belong to quite a new category far transcending all his ancestors in his intellectual characteristics, in his emotional and moral qualities. There is nothing against the assumption, and there is much to be said in its favour, that the last step from animal to man was such an immense one that it brought with it a freedom and [pg 134] richness of psychical life incomparable with anything that had gone before—as if life here realised itself for the first time in very truth, and made everything that previously had been a mere preliminary play.