The first part of the 'Philosophie Zoologique' is the one which deals with the doctrine of evolution or descent with modification. It is to this, therefore, that our attention will be confined. Yet only a comparatively small part of the three hundred and fifty pages which constitute Lamarck's first part are devoted to setting forth the reasons which led him to arrive at his conclusions—the greater part of the volume being occupied with the classification of animals, which we may again omit, as foreign to our purpose.

I shall condense whenever I can, but I do not think the reader will find that I have left out much that bears upon the argument. I shall also use inverted commas while translating with such freedom as to omit several lines together, where I can do so without suppressing anything essential to the elucidation of Lamarck's meaning. I shall, however, throughout refer the reader to the page of the original work from which I am translating.

"The common origin of bodily and mental phenomena," says Lamarck in his preliminary chapter, "has been obscured, because we have studied them chiefly in man, who, as the most highly developed of living beings, presents the problem in its most difficult and complicated aspect. If we had begun our study with that of the lowest organisms, and had proceeded from these to the more complex ones, we should have seen the progression which is observable in organization, and the successive acquisition of various special organs, with new faculties for every additional organ. We should thus have seen that sense of needs—originally hardly perceptible, but gradually increasing in intensity and variety—has led to the attempt to gratify them; that the actions thus induced, having become habitual and energetic, have occasioned the development of organs adapted for their performance; that the force which excites organic movements can in the case of the lowest animals exist outside them and yet animate them; that this force was subsequently introduced into the animals themselves, and fixed within them; and, lastly, that it gave rise to sensibility and, in the end, to intelligence."[208] The reader had better be on his guard here, and whenever Lamarck is speculating about the lowest forms of action and sensation. I have thought it well, however, to give enough of these speculations, as occasion arises, to show their tendency.

"Sensation is not the proximate cause of organic movements. It may be so with the higher animals, but it cannot be shown to be so with plants, nor even with all known animals. At the outset of life there was none of that sensation which could only arise where organic beings had already attained a considerable development. Nature has done all by slow gradations, both organs and faculties being the outcome of a progressive development.[209]

"The mere composition of an animal is but a small part of what deserves study in connection with the animal itself. The effects of its surroundings in causing new wants, the effects of its wants in giving rise to actions, those of its actions in developing habits and tendencies, the effects of use and disuse as affecting any organ, the means which nature takes to preserve and make perfect what has been already acquired—these are all matters of the highest importance.[210]

"In their bearing upon these questions the invertebrate animals are more important and interesting than the vertebrate, for they are more in number, and being more in number are more varied; their variations are more marked, and the steps by which they have advanced in complexity are more easily observed.[211]

"I propose, therefore, to divide this work into three parts, of which the first shall deal with the conventions necessary for the treatment of the subject, the importance of analogical structures, and the meaning which should be attached to the word species. I will point out on the one hand the evidence of a graduated descending scale, as existing between the highest and the lowest organisms; and, on the other, the effect of surroundings and habits on the organs of living beings, as the cause of their development or arrest of development. Lastly, I will treat of the natural order of animals, and show what should be their fittest classification and arrangement."[212]

It seems unnecessary to give Lamarck's intentions with regard to his second and third parts, as they do not here concern us; they deal with the origin of life and mind.

The first chapter of the work opens with the importance of bearing in mind the difference between the conventional and the natural, that is to say, between words and things. Here, as indeed largely throughout this part of his work, he follows Buffon, by whom he is evidently influenced.

"The conventional deals with systems of arrangement, classification, orders, families, genera, and the nomenclature, whether of different sections or of individual objects.